Redescription of male and female genitalia of Tipula(Emodotipula) holoteles(Diptera), with notes on the structure of the semen pump in the subgenus Emodotipula

The original description of male hypopygium of Tipula(Emodotipula) holoteles Alexander, 1924 is expanded with additional morphological characters and detailed illustrations. The additional morphological descriptions of the female ovipositor along with detailed drawings are provided. The male semen pump and the female vaginal apodeme as well as the spermatheca are also described and illustrated for the first time, representing the first report of these structures of Tipula(Emodotipula) holoteles. Additionally, the taxonomic potential of the semen pump in the subgenus Emodotipula is analyzed and discussed. The new specific name Tipula(Emodotipula) yangi nom. nov. is proposed as a replacement name for junior homonym Tipula(Emodotipula) alexanderi Men, 2017. A worldwide checklist and a key to Chinese species of the subgenus Emodotipula are provided.