基金supported by the National Science Foundation of the USA through the grant"Biotic Survey of the Gaoligongshan,a Biodiversity Hotspot in Western Yunnan,China"(No.D EB-0103795)partly supported by the National Natural Sciences Foundation of China(31272271,31272272,30970327)+2 种基金the Foundation of Scientific and Technological Department of Guizhou Province(No.J[2012]2313)the Doctoral Fund of Tongren University(DS1101)the Foundation of Tongren City Scientific and Technological Bureau(No.(2012)63-2)