基金supported by the Program of New Varieties of Genetically Modified Organism Cultivation of China(2009ZX08009-043B,2009ZX08001-006B)‘973’Program(2012CB722504)+2 种基金‘863’Program(2008AA02Z125)National Natural Science Foundation of China for young scholar(31101208,30900263)Zhejiang Provincial Foundation for Natural Science(Y3090665,Y3100573)