基金supported by the Fundamental Research Funds for the Central Non-Profit Research Institution of Chinese Academy of Forestry(CAFYBB2016SY034,key technologies of functional restoration and directional improvement of mudflat wetland)the National Key R&D Program of China(2017YFC0506200)the Research on the synchronicity of dichogamy in Scirpus planiculmis based on population clonal structure(NSFC31800348).