The features of branching and growth studied included height, diameter at breast height (DBH), total number of branches, annual height growth, annual branch elongation in the year of elongating, annual branch number f...The features of branching and growth studied included height, diameter at breast height (DBH), total number of branches, annual height growth, annual branch elongation in the year of elongating, annual branch number for four consecutive years, diameter of branches of different ages, and diameter of stem where branch-whorl originates. For features of total growth and overall branching, no significant differences were found between families, except for DBH. For annual features, no significant differences were found in annual stem height growth, annual branch elongation in the year of elongation and diameter of branches. In the last four years, differences in number of branches were not significant in the first two years but were significant in the last two year; differences in stem diameter where branch-whorls grow were significant for the four consecutive years. Trend of annual growth and branching features of families can be divided into three types as increasing type, stable type and fluctuating type. Most of families have an increasing type with respect of annual height growth and annual branch elongation, while most families belong to a fluctuating type with annual branch number. The results indicated that in the fifth year after planted in seedling seed orchard, differences between families were mostly insignificant. This result may have two main explanations: one is the growth rhyme in early ages of Masson pine, the other one is the complex paternal components to form the open-pollinated families. Family selection seemed to be not useful based on the result. It is suggested to select some of families in the nursery instead of to use all the families when establishing seedling seed orchards with open-pollinated families from plus-trees.展开更多
Through 5 years of phenological observations on Larix principis-rupprechtii Mayr. in primary seed orchard and studies on population and individuals of clones, the annual periodic phenological laws were revealed and th...Through 5 years of phenological observations on Larix principis-rupprechtii Mayr. in primary seed orchard and studies on population and individuals of clones, the annual periodic phenological laws were revealed and the annual phe-nological periodic table was drawn up. The correlation between various phenophases, the air temperature and active accumu-lated temperature were analyzed and expounded. The authors also analyzed the similarities and differences of phenophases among clonal individuals as well as the blooming properties of male and female flowers at the same time. This study could pro-vide theoretical reference for working out the production plan of improved varieties and other management measures in seed orchard of Larix principis-rupprechtii.展开更多
Pinusdensata is one of the main constructive species for coniferous forests in southeast Tibet. P. densata forests are important water conservation forests in the drainage basins of the middle and lower reaches of Yal...Pinusdensata is one of the main constructive species for coniferous forests in southeast Tibet. P. densata forests are important water conservation forests in the drainage basins of the middle and lower reaches of Yalu Tsangpo River, Nyang River and Parlung Zangbo River. In this study, with P. densata forest distributed in southeast Tibet as research object, the seed rain, soil seed bank, seed germination and natural regeneration of P. densata were monitored and ana- lyzed by field investigation, located monitoring and indoor experimental analysis. The results showed that the average intensity of the seed rain of P. densata was 249.30±78.42 seeds/m2, in which the intensity of full seeds was 168.09±56.36 seeds/m2, the intensity of seeds damaged by worms was 41.11±20.25 seeds/m2, and the intensity of empty seeds was 40.10±21.04 seeds/m2. The intensity of the seed rain exhibited a single-peak trend of increasing at first and decreasing then over time. The spatial distribution patterns in the whole seed falling process and at different seed falling time all exhibited clumping distribution, and within in certain range, with the distance from the seed tree increasing, the diffusion intensity of the seed rain was weakened, exhibiting approximately normal distribution. In average density of P. densata seeds in the soil seed bank of P. densata was 231 seeds/m2, in which 62.77% of seeds were distributed in the litter layer, and 37.23% of seeds were distributed in the soil layer, and about 8% of seeds were lost during the pro- cess from seed rain to soil seed bank. Field sowing observation showed that the accumulated germination rate curve of P. densata fitted with Logistics equation y= 91.404/(1+e66194.449). The height structure, basal diameter structure and age structure of seedlings and young trees of P. densata were all of reverse "J" type, indicating good natural regeneration of P. densata. This study would provide a science basis for protection and resource management of P. densata, and further enrich the eval- uation content of national ecological safety curtain of the Tibet plateau.展开更多
The stimulative effect of trace elements on seed germination and seedling growth of Pinus tabulaeformis was tested. The experiments were carried out on seed soak and topdressing with different trace elements and varie...The stimulative effect of trace elements on seed germination and seedling growth of Pinus tabulaeformis was tested. The experiments were carried out on seed soak and topdressing with different trace elements and varied concentrations at the nursery of Gardens Research Institute, Harbin, in 2000-2001. The experimental results showed that soaking seed with 1% and 0.2% concentrations of Mn element produced best result for seed germination, and the germination rate was increased by 9%~19% for the seeds treated with 1% concentration and 12%~14% for the seeds treated with 0.2% concentration compared with the control group. The seeds treated with boron element had lowest germination rate. For trace element topdressing, Mn and Mo elements presented good result for seedling growth and the treatment with low concentration was even better. The height or chlorophyll content of the seedlings with spray of low-concentration Mn and Mo element was much higher than that of untreated ones. In the contrast to the treating method of seed soak, topdressing (application of spraying on foliage) had evident effect on seedling growth.展开更多
Water relation parameters of bare-root seedlings of Chinese fir (Cunninghamia lanceolata Hook.) and Masson pine (Pinus massoniana Lamb.) were measured and changes of root growth potential as well as field survival rat...Water relation parameters of bare-root seedlings of Chinese fir (Cunninghamia lanceolata Hook.) and Masson pine (Pinus massoniana Lamb.) were measured and changes of root growth potential as well as field survival rate of both species were studied after the bare-root seedlings were exposed in a sunny field condition. The results showed that Masson pine had a lower osmotic potential (-2.07Mpa) at turgor loss point and at full turgor (-1.29Mpa), compared with Chinese fir (-1.80Mpa and -1.08Mpa respectively). The parameter Vp/V0 (63.27%) of Masson pine was higher than that of Chinese fir (58.03%). This means that Masson pine has a stronger ability to tolerate desiccation, compared to Chinese fir according to analysis of above water relation parameters. Root growth potential and field survival rate decreased with prolonging duration of exposure. The field survival rate of both species was reduced to less than 40% after the seedling being exposed only two hours. Water poten-tials of 1.60 Mpa and -1.70 Mpa were suggested to be critical values for Chinese fir and Masson pine respectively in successful reforestation.展开更多
35-40-year experience of the study of geographical cultures in Kazakhstan has allowed accumulating of significant scientific information on test of seed progeny of different origins of Pinus sylvestris. Results on gro...35-40-year experience of the study of geographical cultures in Kazakhstan has allowed accumulating of significant scientific information on test of seed progeny of different origins of Pinus sylvestris. Results on growing of climatic types of the pine in provenance trial plantations of Pavlodarskaya, East-Kazakhstan, Kostanayskaya and Akmolinskaya oblasts of the Republic of Kazakhstan are given. Long-term studies have shown that at growing of different origins in determined conditions with the use of identical agricultural techniques, different silvicultural, selection and economic effects are reached. Analysis of all the available material has been conducted on the basis of which new forest seed zoning is designed and the most productive and stable ecotypes are selected as candidates for sort-populations. Arakaragayskiy ecotype is universal for different zones and site conditions of the pine in Kazakhstan. Chebarkulskiy ecotype has good indices on productivity, stability, quality of plantings and on selection effect. Given geographical ecotypes are recommended as candidates for sort-populations.展开更多
Investigation of the above-ground biomass allocation patterns on Scots pine plantations is critical for quantifying the productivity and carbon cycle of forest ecosystems. We estimated above-ground biomass and net pri...Investigation of the above-ground biomass allocation patterns on Scots pine plantations is critical for quantifying the productivity and carbon cycle of forest ecosystems. We estimated above-ground biomass and net primary production of a 25-year-old Pinus sylvestris L. (Scots pine) plantation, in a semi-arid region of Mongolia. The above-ground biomass of sample trees was divided into stem wood, stem bark, live branches, dead branches and needles. Total biomass for the stand was only 18.03 Mg ha1, of which 47.6% was found in stem wood, 25.8% in live branches and 14.8% in needles. The growth rate of the Scots pine plantation in the study region was relatively low compared with other regions. In the study area, it was observed that the rate of biomass accumulation in the plantation was very slow; this can be explained by very limited growing conditions and intensive crown closure. The results from this study indicate that it may be necessary to carry out thinning to increase biomass production by reducing competition between trees in the Scotch pine plantation.展开更多
Pollination dynamics highly determines the genetic quality of seed orchard crops. However, there is less research about the effect of mating patterns on seed productivity of orchard crops. So far, clonal seed orchards...Pollination dynamics highly determines the genetic quality of seed orchard crops. However, there is less research about the effect of mating patterns on seed productivity of orchard crops. So far, clonal seed orchards have been producing genetically improved seedlings used for most Japanese larch(Larix kaempferi(Lamb.) Carr.) plantations in China. In the present study, a total of 17 highly variable simple sequence repeat(SSR) markers were used for genotyping a progeny trial population consisting of 647 open-pollinated progenies germinated from seeds which were collected from 63 maternal clones with 140 potential paternal clones in a Japanese larch clonal seed orchard in China. Paternity analysis was used in the present case study in order to evaluate the level of paternal gametic contribution, estimate pollen contamination and selfing rates, and investigate pollination patterns,pollen dispersal patterns and the impact of mating patterns on seed productivity of orchard crops. We observed 93.7% of the success rate of the parental assignment, unequal paternal gametic contribution(0–12.4%) with 6.3% of the progenies derived from pollen contamination or unsampled pollen donors, and absence of evidence for selfing. We also found that pollination rate highly depended on the distance between pollen donors and maternal parents, the majority of the identified crossing(65.7%)occurred between clones within a 150-m radius, and large variations in growth performance existed among the paternal halfsiblings. Progeny growth performance(diameter at breast(DBH) and height(HGT)) was measured at Age-20 in order to investigate the impact of mating patterns on timber production of orchard crops. As either the paternal or maternal, two clones(i.e., clones Z38 and Z62) were identified to have produced progenies with higher average stem volume breeding values than that of all of the progenies. Specifically, the genetic gains for volume were 3.53% for the two clones as paternal parents, and 8.26% as the maternal parents at Age-20. Thus, both elite clones were ideal candidates for the construction of next-generation clonal seed orchards due to their synchronous reproductive phenology with greater crossing rate and higher genetic gain. These results improved the pedigree information to provide solid evidence of mating patterns for future design and effective management of seed orchards and for the development of viable long-term breeding strategies for other coniferous species.展开更多
基金This paper was a part of the National Key Project of Science and Technology on Masson Pine breeding during 1996-2000.
文摘The features of branching and growth studied included height, diameter at breast height (DBH), total number of branches, annual height growth, annual branch elongation in the year of elongating, annual branch number for four consecutive years, diameter of branches of different ages, and diameter of stem where branch-whorl originates. For features of total growth and overall branching, no significant differences were found between families, except for DBH. For annual features, no significant differences were found in annual stem height growth, annual branch elongation in the year of elongation and diameter of branches. In the last four years, differences in number of branches were not significant in the first two years but were significant in the last two year; differences in stem diameter where branch-whorls grow were significant for the four consecutive years. Trend of annual growth and branching features of families can be divided into three types as increasing type, stable type and fluctuating type. Most of families have an increasing type with respect of annual height growth and annual branch elongation, while most families belong to a fluctuating type with annual branch number. The results indicated that in the fifth year after planted in seedling seed orchard, differences between families were mostly insignificant. This result may have two main explanations: one is the growth rhyme in early ages of Masson pine, the other one is the complex paternal components to form the open-pollinated families. Family selection seemed to be not useful based on the result. It is suggested to select some of families in the nursery instead of to use all the families when establishing seedling seed orchards with open-pollinated families from plus-trees.
文摘Through 5 years of phenological observations on Larix principis-rupprechtii Mayr. in primary seed orchard and studies on population and individuals of clones, the annual periodic phenological laws were revealed and the annual phe-nological periodic table was drawn up. The correlation between various phenophases, the air temperature and active accumu-lated temperature were analyzed and expounded. The authors also analyzed the similarities and differences of phenophases among clonal individuals as well as the blooming properties of male and female flowers at the same time. This study could pro-vide theoretical reference for working out the production plan of improved varieties and other management measures in seed orchard of Larix principis-rupprechtii.
基金Supported by Natural Science Foundation of China(31460200)Natural Science Foundation of Tibet Autonomous Region(2016ZR-15-40)~~
文摘Pinusdensata is one of the main constructive species for coniferous forests in southeast Tibet. P. densata forests are important water conservation forests in the drainage basins of the middle and lower reaches of Yalu Tsangpo River, Nyang River and Parlung Zangbo River. In this study, with P. densata forest distributed in southeast Tibet as research object, the seed rain, soil seed bank, seed germination and natural regeneration of P. densata were monitored and ana- lyzed by field investigation, located monitoring and indoor experimental analysis. The results showed that the average intensity of the seed rain of P. densata was 249.30±78.42 seeds/m2, in which the intensity of full seeds was 168.09±56.36 seeds/m2, the intensity of seeds damaged by worms was 41.11±20.25 seeds/m2, and the intensity of empty seeds was 40.10±21.04 seeds/m2. The intensity of the seed rain exhibited a single-peak trend of increasing at first and decreasing then over time. The spatial distribution patterns in the whole seed falling process and at different seed falling time all exhibited clumping distribution, and within in certain range, with the distance from the seed tree increasing, the diffusion intensity of the seed rain was weakened, exhibiting approximately normal distribution. In average density of P. densata seeds in the soil seed bank of P. densata was 231 seeds/m2, in which 62.77% of seeds were distributed in the litter layer, and 37.23% of seeds were distributed in the soil layer, and about 8% of seeds were lost during the pro- cess from seed rain to soil seed bank. Field sowing observation showed that the accumulated germination rate curve of P. densata fitted with Logistics equation y= 91.404/(1+e66194.449). The height structure, basal diameter structure and age structure of seedlings and young trees of P. densata were all of reverse "J" type, indicating good natural regeneration of P. densata. This study would provide a science basis for protection and resource management of P. densata, and further enrich the eval- uation content of national ecological safety curtain of the Tibet plateau.
文摘The stimulative effect of trace elements on seed germination and seedling growth of Pinus tabulaeformis was tested. The experiments were carried out on seed soak and topdressing with different trace elements and varied concentrations at the nursery of Gardens Research Institute, Harbin, in 2000-2001. The experimental results showed that soaking seed with 1% and 0.2% concentrations of Mn element produced best result for seed germination, and the germination rate was increased by 9%~19% for the seeds treated with 1% concentration and 12%~14% for the seeds treated with 0.2% concentration compared with the control group. The seeds treated with boron element had lowest germination rate. For trace element topdressing, Mn and Mo elements presented good result for seedling growth and the treatment with low concentration was even better. The height or chlorophyll content of the seedlings with spray of low-concentration Mn and Mo element was much higher than that of untreated ones. In the contrast to the treating method of seed soak, topdressing (application of spraying on foliage) had evident effect on seedling growth.
基金This research was supported by the Sciences Research Foundation of Nanjing Forestry University.
文摘Water relation parameters of bare-root seedlings of Chinese fir (Cunninghamia lanceolata Hook.) and Masson pine (Pinus massoniana Lamb.) were measured and changes of root growth potential as well as field survival rate of both species were studied after the bare-root seedlings were exposed in a sunny field condition. The results showed that Masson pine had a lower osmotic potential (-2.07Mpa) at turgor loss point and at full turgor (-1.29Mpa), compared with Chinese fir (-1.80Mpa and -1.08Mpa respectively). The parameter Vp/V0 (63.27%) of Masson pine was higher than that of Chinese fir (58.03%). This means that Masson pine has a stronger ability to tolerate desiccation, compared to Chinese fir according to analysis of above water relation parameters. Root growth potential and field survival rate decreased with prolonging duration of exposure. The field survival rate of both species was reduced to less than 40% after the seedling being exposed only two hours. Water poten-tials of 1.60 Mpa and -1.70 Mpa were suggested to be critical values for Chinese fir and Masson pine respectively in successful reforestation.
文摘35-40-year experience of the study of geographical cultures in Kazakhstan has allowed accumulating of significant scientific information on test of seed progeny of different origins of Pinus sylvestris. Results on growing of climatic types of the pine in provenance trial plantations of Pavlodarskaya, East-Kazakhstan, Kostanayskaya and Akmolinskaya oblasts of the Republic of Kazakhstan are given. Long-term studies have shown that at growing of different origins in determined conditions with the use of identical agricultural techniques, different silvicultural, selection and economic effects are reached. Analysis of all the available material has been conducted on the basis of which new forest seed zoning is designed and the most productive and stable ecotypes are selected as candidates for sort-populations. Arakaragayskiy ecotype is universal for different zones and site conditions of the pine in Kazakhstan. Chebarkulskiy ecotype has good indices on productivity, stability, quality of plantings and on selection effect. Given geographical ecotypes are recommended as candidates for sort-populations.
文摘Investigation of the above-ground biomass allocation patterns on Scots pine plantations is critical for quantifying the productivity and carbon cycle of forest ecosystems. We estimated above-ground biomass and net primary production of a 25-year-old Pinus sylvestris L. (Scots pine) plantation, in a semi-arid region of Mongolia. The above-ground biomass of sample trees was divided into stem wood, stem bark, live branches, dead branches and needles. Total biomass for the stand was only 18.03 Mg ha1, of which 47.6% was found in stem wood, 25.8% in live branches and 14.8% in needles. The growth rate of the Scots pine plantation in the study region was relatively low compared with other regions. In the study area, it was observed that the rate of biomass accumulation in the plantation was very slow; this can be explained by very limited growing conditions and intensive crown closure. The results from this study indicate that it may be necessary to carry out thinning to increase biomass production by reducing competition between trees in the Scotch pine plantation.
基金supported by the Forestry Industry Research Special Funds for Public Welfare Projects (201504104)
文摘Pollination dynamics highly determines the genetic quality of seed orchard crops. However, there is less research about the effect of mating patterns on seed productivity of orchard crops. So far, clonal seed orchards have been producing genetically improved seedlings used for most Japanese larch(Larix kaempferi(Lamb.) Carr.) plantations in China. In the present study, a total of 17 highly variable simple sequence repeat(SSR) markers were used for genotyping a progeny trial population consisting of 647 open-pollinated progenies germinated from seeds which were collected from 63 maternal clones with 140 potential paternal clones in a Japanese larch clonal seed orchard in China. Paternity analysis was used in the present case study in order to evaluate the level of paternal gametic contribution, estimate pollen contamination and selfing rates, and investigate pollination patterns,pollen dispersal patterns and the impact of mating patterns on seed productivity of orchard crops. We observed 93.7% of the success rate of the parental assignment, unequal paternal gametic contribution(0–12.4%) with 6.3% of the progenies derived from pollen contamination or unsampled pollen donors, and absence of evidence for selfing. We also found that pollination rate highly depended on the distance between pollen donors and maternal parents, the majority of the identified crossing(65.7%)occurred between clones within a 150-m radius, and large variations in growth performance existed among the paternal halfsiblings. Progeny growth performance(diameter at breast(DBH) and height(HGT)) was measured at Age-20 in order to investigate the impact of mating patterns on timber production of orchard crops. As either the paternal or maternal, two clones(i.e., clones Z38 and Z62) were identified to have produced progenies with higher average stem volume breeding values than that of all of the progenies. Specifically, the genetic gains for volume were 3.53% for the two clones as paternal parents, and 8.26% as the maternal parents at Age-20. Thus, both elite clones were ideal candidates for the construction of next-generation clonal seed orchards due to their synchronous reproductive phenology with greater crossing rate and higher genetic gain. These results improved the pedigree information to provide solid evidence of mating patterns for future design and effective management of seed orchards and for the development of viable long-term breeding strategies for other coniferous species.