We propose a solvable aggregation model to mimic the evolution of population A, asset B, and the quantifiable resource C in a society. In this system, the population and asset aggregates themselves grow through selfex...We propose a solvable aggregation model to mimic the evolution of population A, asset B, and the quantifiable resource C in a society. In this system, the population and asset aggregates themselves grow through selfexchanges with the rate kernels Kl(k,j) = K1kj and K2(h,j) = K2kj, respectively. The actions of the population and asset aggregations on the aggregation evolution of resource aggregates are described by the population-catalyzed monomer death of resource aggregates and asset-catalyzed monomer birth of resource aggregates with the rate kerne/s J1(k,j)=J1k and J2(k,j) = J2k, respectively. Meanwhile, the asset and resource aggregates conjunctly catalyze the monomer birth of population aggregates with the rate kernel I1 (k,i,j) = I1ki^μjη, and population and resource aggregates conjunctly catalyze the monomer birth of asset aggregates with the rate kernel /2(k, i, j) = I2ki^νj^η. The kinetic behaviors of species A, B, and C are investigated by means of the mean-field rate equation approach. The effects of the population-catalyzed death and asset-catalyzed birth on the evolution of resource aggregates based on the self-exchanges of population and asset appear in effective forms. The coefficients of the effective population-catalyzed death and the asset-catalyzed birth are expressed as J1e = J1/K1 and J2e= J2/K2, respectively. The aggregate size distribution of C species is found to be crucially dominated by the competition between the effective death and the effective birth. It satisfies the conventional scaling form, generalized scaling form, and modified scaling form in the cases of J1e〈J2e, J1e=J2e, and J1e〉J2e, respectively. Meanwhile, we also find the aggregate size distributions of populations and assets both fall into two distinct categories for different parameters μ,ν, and η: (i) When μ=ν=η=0 and μ=ν=η=1, the population and asset aggregates obey the generalized scaling forms; and (ii) When μ=ν=1,η=0, and μ=ν=η=1, the population and asset aggregates experience gelation transitions at finite times and the scaling forms break down.展开更多
Unlike most brood parasites, several species of cowbird (Molothrus) are generalists that parasitize multiple host species across their range and within the same communities; likewise, there is little evidence that ind...Unlike most brood parasites, several species of cowbird (Molothrus) are generalists that parasitize multiple host species across their range and within the same communities; likewise, there is little evidence that individuals within a population specialize on host species. This situation has variously been attributed to the recency of cowbird evolution (the 'evolutionary lag' hypothesis) or to hidden costs of rejection by hosts (the 'equilibrium' hypothesis). Both hypotheses have some support as cowbirds are indeed a relatively young clade compared with more specialized cuckoos and cowbirds are capable of sophisticated behaviors such as retaliation against rejection ('mafia' behaviors) that would select for acceptance of cowbird eggs. Nevertheless, many species in the Americas have evolved specialized defenses against cowbird parasitism, almost all of which live in more open habitats (e. g., grasslands, shrublands, riparian strips), which indicates that coevolutionary processes can operate in ways that select for host defenses in spite of cowbird counterdefenses. We propose that the structure of landscapes in North America may explain why forest-nesting birds lack defenses against parasitism and reinforce the long-term maintenance of generalized brood parasitism in cowbirds. Because cowbirds require open habitats in which to feed, they are rare or absent in large forest tracts, which dominate much of the forest cover of the Americas. These tracts act as 'source habitats' that produce surplus young that recolonize populations in smaller, fragmented forest patches in which rates of both cowbird parasitism and nest predation are very high ('sink' habitats). Evolution of antiparasite adaptations would be very slow in this situation because most hosts are produced in areas where there is little or no cowbird parasitism. In addition, the interplay of host breeding dispersal, source-sink metapopulation dynamics, and fragmented forest habitat could further deter the evolution of host defenses against parasitism. Therefore, as long as large forest tracts remain widespread in North America, most forest birds will likely continue to lack defenses against cowbird parasitism, guaranteeing a steady supply of na ve hosts in forest habitats, even in fragmented landscapes. This situation will, in turn, favor host generalist cowbirds that actively avoid more open habitats in favor of parasitizing forest bird communities. These forest communities may also act as source populations for cowbirds, which might pump surplus generalist cowbirds into more open habitats further slowing the coevolutionary process. As long as large forest tracts are a common part of the landscape, generalist parasitism may persist indefinitely.展开更多
The exploitation of renewable resources creates many complex problems for culture, ecology and economics as well. Ascertaining the essentials behind the coraplex problems is very important, in this paper, we mainly st...The exploitation of renewable resources creates many complex problems for culture, ecology and economics as well. Ascertaining the essentials behind the coraplex problems is very important, in this paper, we mainly study various complex relations appearing in the optimal exploitation process for renewable resources. First, we derive a sufficient condition on the existence of optimal harvesting policies for one-species population resources. Then we present every possible optimal harvesting pattern for such a modeh On the basis of this, we give a computing formula for estimating the optimal harvesting period, optimal transitional period, and optimal recruitment period. The main difference with respect to the previous works in literature is that our optimal harvesting policy is a piece-wise continuous function of time t, at the piecewise point to, which is called switching time. At the switching time we switch the harvesting rate from h to some transitional control u,, then to 0. Clearly this kind of harvesting policy is easier to carry out than those by others, provided that there exists a managing department which can highly supervise the resources.展开更多
Many territorial animals exhibit reduced aggression towards neighbours. Known as "the dear enemy effect", this phenomenon has been documented among conspecific animals across a wide range of animal taxa. In theory, ...Many territorial animals exhibit reduced aggression towards neighbours. Known as "the dear enemy effect", this phenomenon has been documented among conspecific animals across a wide range of animal taxa. In theory, the dear enemy ef- fect can also exist between individuals of different species, particularly when those species compete for shared resources. To date, a heterospecific dear enemy effects has only been documented in ants. In this study, we test for both a conspecific and heterospe- cific dear enemy effect in neotropical rufous-and-white wrens Thryophilus rufalbus. This species competes for resources with banded wrens Thryophilus pleurostictus, a closely related sympatric congener. We used acoustic playback to simulate ruf- ous-and-white wren and banded wren neighbours and non-neighbours at the edges of rufous-and-white wren territories. Ruf- ous-and-white wrens responded more strongly to signals from their own species, demonstrating that resident males discriminate between conspecific and heterospecific rivals. They did not, however, exhibit a conspecific dear enemy effect. Further, they did not exhibit a heterospecific dear enemy effect. This could be due to neighbours and non-neighbours posing similar levels of threat in this system, to the possibility that playback from the edges of the subjects' large territories did not simulate a threatening signal, or to other factors. Our study provides the first test of a heterospecific dear enemy effect in vertebrates, and presents a valuable experimental approach for testing for a heterospecific dear enemy effect in other animals [Current Zoology 61 (1): 23-33, 2015].展开更多
In this paper the management model a two-species fishery involving impulsesis investigated by using optimal impulsive control theorem. Optimal impulsive harvesting times andthe corresponding optimal harvesting populat...In this paper the management model a two-species fishery involving impulsesis investigated by using optimal impulsive control theorem. Optimal impulsive harvesting times andthe corresponding optimal harvesting population levels in different cases are obtained.展开更多
Aims Long-term heavy grazing reduces plant diversity and ecosystem function by intensifying nitrogen(N)and water limitation.In contrast,the absence of biomass removal can cause species loss by elevating light competit...Aims Long-term heavy grazing reduces plant diversity and ecosystem function by intensifying nitrogen(N)and water limitation.In contrast,the absence of biomass removal can cause species loss by elevating light competition and weakening community stability,which is exacerbated by N and water enrichment.Hence,how to maintain species diversity and community stability is still a huge challenge for sustainable management of worldwide grasslands.Methods We conducted a 4-year manipulated experiment in six long-term grazing blocks to explore combination of resource additions and biomass removal(increased water,N and light availability)on species richness and community stability in semiarid grasslands of Inner Mongolia,China.Important Findings In all blocks treated with the combination of resource additions and biomass removal,primary productivity increased and species richness and community stability were maintained over 4 years of experiment.At both species and plant functional group(PFG)levels,the aboveground biomass of treated plants remained temporally stable in treatments with the combination of N and/or water addition and biomass removal.The maintenance of species richness was primarily caused by the biomass removal,which could increase the amount of light exposure for grasses under resource enrichment.Both species asynchrony and stability of PFGs contributed to the high temporal stability observed in these communities.Our results indicate that management practices of combined resource enrichment with biomass removal,such as grazing or mowing,could not only enhance primary productivity but also maintain plant species diversity,species asynchrony and community stability.Furthermore,as overgrazing-induced degradation and resource enrichment-induced biodiversity loss continue to be major problems worldwide,our findings have important implications for adaptive management in semiarid grasslands and beyond.展开更多
In order to understand the pattern and status of plant species in the forest-meadow ecotonein Sanjiangyuan Nature Reserve, this study was undertaken through expert consultation, literaturesurvey and fieldwork in vario...In order to understand the pattern and status of plant species in the forest-meadow ecotonein Sanjiangyuan Nature Reserve, this study was undertaken through expert consultation, literaturesurvey and fieldwork in various parts of the ecotone during 2002-2004. Based on 3 important systems,i.e. threatened status, genetic loss effect and use value, the list of threatened and prioritized plants wasestablished. In this paper, the threatened coefficient was defined as the criteria of category for threatenedplants, the conservation priorities coefficient summed by threatened coefficient, genetic coefficient anduse value coefficient was defined as the criteria of prioritized plants. In addition, 10 indicators toevaluate threatened grading and conservation priorities sequence were included in the 3 importantsystems. The weights of 3 systems and 10 indicators were given through expert consultation andanalytic hierarchy process. Ten indicators were given scores based on the subcriteria, respectively, andall the scores of the 3 systems were summed up for each species, then contrasted to the criteria ofthreatened grading and conservation priorities for plant species. The status of endangerment andconservation priorities of plant species were analyzed, and the results showed that the number ofendangered species was 2, vulnerable species 37, near threatened species 140, safe species 767; the firstconcern category had 4 species, the second concern category 18, the third concern category 150 and theleast concern category 774.展开更多
基金Supported by the National Natural Science Foundation of China under Grant Nos. 10775104, 10275048, and 10305009the Zhejiang Provincial Natural Science Foundation of China under Grant No. 102067
文摘We propose a solvable aggregation model to mimic the evolution of population A, asset B, and the quantifiable resource C in a society. In this system, the population and asset aggregates themselves grow through selfexchanges with the rate kernels Kl(k,j) = K1kj and K2(h,j) = K2kj, respectively. The actions of the population and asset aggregations on the aggregation evolution of resource aggregates are described by the population-catalyzed monomer death of resource aggregates and asset-catalyzed monomer birth of resource aggregates with the rate kerne/s J1(k,j)=J1k and J2(k,j) = J2k, respectively. Meanwhile, the asset and resource aggregates conjunctly catalyze the monomer birth of population aggregates with the rate kernel I1 (k,i,j) = I1ki^μjη, and population and resource aggregates conjunctly catalyze the monomer birth of asset aggregates with the rate kernel /2(k, i, j) = I2ki^νj^η. The kinetic behaviors of species A, B, and C are investigated by means of the mean-field rate equation approach. The effects of the population-catalyzed death and asset-catalyzed birth on the evolution of resource aggregates based on the self-exchanges of population and asset appear in effective forms. The coefficients of the effective population-catalyzed death and the asset-catalyzed birth are expressed as J1e = J1/K1 and J2e= J2/K2, respectively. The aggregate size distribution of C species is found to be crucially dominated by the competition between the effective death and the effective birth. It satisfies the conventional scaling form, generalized scaling form, and modified scaling form in the cases of J1e〈J2e, J1e=J2e, and J1e〉J2e, respectively. Meanwhile, we also find the aggregate size distributions of populations and assets both fall into two distinct categories for different parameters μ,ν, and η: (i) When μ=ν=η=0 and μ=ν=η=1, the population and asset aggregates obey the generalized scaling forms; and (ii) When μ=ν=1,η=0, and μ=ν=η=1, the population and asset aggregates experience gelation transitions at finite times and the scaling forms break down.
文摘Unlike most brood parasites, several species of cowbird (Molothrus) are generalists that parasitize multiple host species across their range and within the same communities; likewise, there is little evidence that individuals within a population specialize on host species. This situation has variously been attributed to the recency of cowbird evolution (the 'evolutionary lag' hypothesis) or to hidden costs of rejection by hosts (the 'equilibrium' hypothesis). Both hypotheses have some support as cowbirds are indeed a relatively young clade compared with more specialized cuckoos and cowbirds are capable of sophisticated behaviors such as retaliation against rejection ('mafia' behaviors) that would select for acceptance of cowbird eggs. Nevertheless, many species in the Americas have evolved specialized defenses against cowbird parasitism, almost all of which live in more open habitats (e. g., grasslands, shrublands, riparian strips), which indicates that coevolutionary processes can operate in ways that select for host defenses in spite of cowbird counterdefenses. We propose that the structure of landscapes in North America may explain why forest-nesting birds lack defenses against parasitism and reinforce the long-term maintenance of generalized brood parasitism in cowbirds. Because cowbirds require open habitats in which to feed, they are rare or absent in large forest tracts, which dominate much of the forest cover of the Americas. These tracts act as 'source habitats' that produce surplus young that recolonize populations in smaller, fragmented forest patches in which rates of both cowbird parasitism and nest predation are very high ('sink' habitats). Evolution of antiparasite adaptations would be very slow in this situation because most hosts are produced in areas where there is little or no cowbird parasitism. In addition, the interplay of host breeding dispersal, source-sink metapopulation dynamics, and fragmented forest habitat could further deter the evolution of host defenses against parasitism. Therefore, as long as large forest tracts remain widespread in North America, most forest birds will likely continue to lack defenses against cowbird parasitism, guaranteeing a steady supply of na ve hosts in forest habitats, even in fragmented landscapes. This situation will, in turn, favor host generalist cowbirds that actively avoid more open habitats in favor of parasitizing forest bird communities. These forest communities may also act as source populations for cowbirds, which might pump surplus generalist cowbirds into more open habitats further slowing the coevolutionary process. As long as large forest tracts are a common part of the landscape, generalist parasitism may persist indefinitely.
基金This Work is supported by National Natural Science Fund of China Under Grant No.70271066
文摘The exploitation of renewable resources creates many complex problems for culture, ecology and economics as well. Ascertaining the essentials behind the coraplex problems is very important, in this paper, we mainly study various complex relations appearing in the optimal exploitation process for renewable resources. First, we derive a sufficient condition on the existence of optimal harvesting policies for one-species population resources. Then we present every possible optimal harvesting pattern for such a modeh On the basis of this, we give a computing formula for estimating the optimal harvesting period, optimal transitional period, and optimal recruitment period. The main difference with respect to the previous works in literature is that our optimal harvesting policy is a piece-wise continuous function of time t, at the piecewise point to, which is called switching time. At the switching time we switch the harvesting rate from h to some transitional control u,, then to 0. Clearly this kind of harvesting policy is easier to carry out than those by others, provided that there exists a managing department which can highly supervise the resources.
文摘Many territorial animals exhibit reduced aggression towards neighbours. Known as "the dear enemy effect", this phenomenon has been documented among conspecific animals across a wide range of animal taxa. In theory, the dear enemy ef- fect can also exist between individuals of different species, particularly when those species compete for shared resources. To date, a heterospecific dear enemy effects has only been documented in ants. In this study, we test for both a conspecific and heterospe- cific dear enemy effect in neotropical rufous-and-white wrens Thryophilus rufalbus. This species competes for resources with banded wrens Thryophilus pleurostictus, a closely related sympatric congener. We used acoustic playback to simulate ruf- ous-and-white wren and banded wren neighbours and non-neighbours at the edges of rufous-and-white wren territories. Ruf- ous-and-white wrens responded more strongly to signals from their own species, demonstrating that resident males discriminate between conspecific and heterospecific rivals. They did not, however, exhibit a conspecific dear enemy effect. Further, they did not exhibit a heterospecific dear enemy effect. This could be due to neighbours and non-neighbours posing similar levels of threat in this system, to the possibility that playback from the edges of the subjects' large territories did not simulate a threatening signal, or to other factors. Our study provides the first test of a heterospecific dear enemy effect in vertebrates, and presents a valuable experimental approach for testing for a heterospecific dear enemy effect in other animals [Current Zoology 61 (1): 23-33, 2015].
基金This research is supported by the Foundation of Educational Committee of Liaoning Province (No.20331080)
文摘In this paper the management model a two-species fishery involving impulsesis investigated by using optimal impulsive control theorem. Optimal impulsive harvesting times andthe corresponding optimal harvesting population levels in different cases are obtained.
基金supported by grants from the National Natural Science Foundation of China(31630010 and 31320103916).
文摘Aims Long-term heavy grazing reduces plant diversity and ecosystem function by intensifying nitrogen(N)and water limitation.In contrast,the absence of biomass removal can cause species loss by elevating light competition and weakening community stability,which is exacerbated by N and water enrichment.Hence,how to maintain species diversity and community stability is still a huge challenge for sustainable management of worldwide grasslands.Methods We conducted a 4-year manipulated experiment in six long-term grazing blocks to explore combination of resource additions and biomass removal(increased water,N and light availability)on species richness and community stability in semiarid grasslands of Inner Mongolia,China.Important Findings In all blocks treated with the combination of resource additions and biomass removal,primary productivity increased and species richness and community stability were maintained over 4 years of experiment.At both species and plant functional group(PFG)levels,the aboveground biomass of treated plants remained temporally stable in treatments with the combination of N and/or water addition and biomass removal.The maintenance of species richness was primarily caused by the biomass removal,which could increase the amount of light exposure for grasses under resource enrichment.Both species asynchrony and stability of PFGs contributed to the high temporal stability observed in these communities.Our results indicate that management practices of combined resource enrichment with biomass removal,such as grazing or mowing,could not only enhance primary productivity but also maintain plant species diversity,species asynchrony and community stability.Furthermore,as overgrazing-induced degradation and resource enrichment-induced biodiversity loss continue to be major problems worldwide,our findings have important implications for adaptive management in semiarid grasslands and beyond.
基金This research is one part of "The Rapid Evaluation Technology of Threatened Species in Sanjiangyuan Nature Reserve " (2001BA510B10-03), which is supported by the National Advanced Project of the Tenth Five-year Plan.
文摘In order to understand the pattern and status of plant species in the forest-meadow ecotonein Sanjiangyuan Nature Reserve, this study was undertaken through expert consultation, literaturesurvey and fieldwork in various parts of the ecotone during 2002-2004. Based on 3 important systems,i.e. threatened status, genetic loss effect and use value, the list of threatened and prioritized plants wasestablished. In this paper, the threatened coefficient was defined as the criteria of category for threatenedplants, the conservation priorities coefficient summed by threatened coefficient, genetic coefficient anduse value coefficient was defined as the criteria of prioritized plants. In addition, 10 indicators toevaluate threatened grading and conservation priorities sequence were included in the 3 importantsystems. The weights of 3 systems and 10 indicators were given through expert consultation andanalytic hierarchy process. Ten indicators were given scores based on the subcriteria, respectively, andall the scores of the 3 systems were summed up for each species, then contrasted to the criteria ofthreatened grading and conservation priorities for plant species. The status of endangerment andconservation priorities of plant species were analyzed, and the results showed that the number ofendangered species was 2, vulnerable species 37, near threatened species 140, safe species 767; the firstconcern category had 4 species, the second concern category 18, the third concern category 150 and theleast concern category 774.