The series Variantia Ching et S. H. Wu mainly occur in China and its members are highly variable in morphology. The denomination on this group of Asplenium is very confused in the herbaria. We hop e by means of a bios...The series Variantia Ching et S. H. Wu mainly occur in China and its members are highly variable in morphology. The denomination on this group of Asplenium is very confused in the herbaria. We hop e by means of a biosystematic study to find out their genetic relationships in the reticulate evolution, and to raise a suggestion on their taxonomic treatment. Evidence from cytology, allozyme, morphology, and palynology shows that three ancestor diploids have formed Asplenium sarelii complex comprising 13 members. A. sarelii Hook. should be typified as a diploid. The so-called tetraploid 'A. sarelii' before is an allotetraploid that comes from the doubled hybrid between diploid A. sarelii and A. tenuicaule Hayata, which should be treated as a new species A. wudangense Z. R. Wang et X. Hou. A. pekinense Hance is an autotetraploid that comes from the doubled diploid ancestor A. sarelii. A. lushanense C. Chr., a diploid species and the only ancestor of A. yunnariense group, should not been sunk as a synonym of tetraploid A. yunnariense Franch. Most probably, A. varians Wall. ex Hook. et Grev. is an autotetraploid of A. tenuicaule Hayata. Three new natural tetraploid hybrids and their origins have been found out: they are A. x longmenense ( = A. pekinense x varians), A. x jingyunense ( = A. pekinense x yunnanense) and A. x kidoi ( = A. pekinense x wudangense). Three other new natural triploid hybrids have been found and their origins have been inferred: they are A. X huawuense ( = A. sarelii X wudangense), A. x luyunense ( = A. lushanense x yunnanense) and A. x teniuvaians ( = A. tenuicaule x varians). The method of allozyme comparion combined with cytological observation is employed to reveal the complicated relationships among the members of Asplenium sarelii complex in reticulate evolution and proved to be a highly effective tool to investigate the origin of polyploid and hybrid.展开更多
The Lipovtsy flora originates mostly from the synonymous Formation exposed at the coal quarries in the Razdolnaya( Suifun) River Basin in Heilongjiang Province( China) and Primorye Region( Russia). The thorough study ...The Lipovtsy flora originates mostly from the synonymous Formation exposed at the coal quarries in the Razdolnaya( Suifun) River Basin in Heilongjiang Province( China) and Primorye Region( Russia). The thorough study of this flora has revealed its high diversity and abundance. One of the most prominent groups in the Lipovtsy flora is the ferns. Next in diversity are conifers( Miroviaceae,Taxodiaceae,Podozamitaceae,Taxaceae,Cephalotaxaceae). It was revealed the remarkable quantity of bennettitalean remains( 24 taxa). The main peculiarity of this flora is the appearance of first angiosperms( pollen,dispersed cuticle,leaves). The fine preservation of the cuticles from the coals allows us to research the taxonomical composition of peat-forming plants and clear main components of the swamp plant communities. These plants supplied the material for the accumulation of resinite coals on the territory of Razdolnaya River Basin were conifers Miroviaceae,taxodialeans,bennettites,as well as gleicheniaceous and cyatheaceous ferns.展开更多
文摘The series Variantia Ching et S. H. Wu mainly occur in China and its members are highly variable in morphology. The denomination on this group of Asplenium is very confused in the herbaria. We hop e by means of a biosystematic study to find out their genetic relationships in the reticulate evolution, and to raise a suggestion on their taxonomic treatment. Evidence from cytology, allozyme, morphology, and palynology shows that three ancestor diploids have formed Asplenium sarelii complex comprising 13 members. A. sarelii Hook. should be typified as a diploid. The so-called tetraploid 'A. sarelii' before is an allotetraploid that comes from the doubled hybrid between diploid A. sarelii and A. tenuicaule Hayata, which should be treated as a new species A. wudangense Z. R. Wang et X. Hou. A. pekinense Hance is an autotetraploid that comes from the doubled diploid ancestor A. sarelii. A. lushanense C. Chr., a diploid species and the only ancestor of A. yunnariense group, should not been sunk as a synonym of tetraploid A. yunnariense Franch. Most probably, A. varians Wall. ex Hook. et Grev. is an autotetraploid of A. tenuicaule Hayata. Three new natural tetraploid hybrids and their origins have been found out: they are A. x longmenense ( = A. pekinense x varians), A. x jingyunense ( = A. pekinense x yunnanense) and A. x kidoi ( = A. pekinense x wudangense). Three other new natural triploid hybrids have been found and their origins have been inferred: they are A. X huawuense ( = A. sarelii X wudangense), A. x luyunense ( = A. lushanense x yunnanense) and A. x teniuvaians ( = A. tenuicaule x varians). The method of allozyme comparion combined with cytological observation is employed to reveal the complicated relationships among the members of Asplenium sarelii complex in reticulate evolution and proved to be a highly effective tool to investigate the origin of polyploid and hybrid.
基金Supported by project of Russian Foundation for Basic Research(No.16-04-01411)
文摘The Lipovtsy flora originates mostly from the synonymous Formation exposed at the coal quarries in the Razdolnaya( Suifun) River Basin in Heilongjiang Province( China) and Primorye Region( Russia). The thorough study of this flora has revealed its high diversity and abundance. One of the most prominent groups in the Lipovtsy flora is the ferns. Next in diversity are conifers( Miroviaceae,Taxodiaceae,Podozamitaceae,Taxaceae,Cephalotaxaceae). It was revealed the remarkable quantity of bennettitalean remains( 24 taxa). The main peculiarity of this flora is the appearance of first angiosperms( pollen,dispersed cuticle,leaves). The fine preservation of the cuticles from the coals allows us to research the taxonomical composition of peat-forming plants and clear main components of the swamp plant communities. These plants supplied the material for the accumulation of resinite coals on the territory of Razdolnaya River Basin were conifers Miroviaceae,taxodialeans,bennettites,as well as gleicheniaceous and cyatheaceous ferns.
