基金Supported by National Key Technology Research and Development Program of China(2011BAD16B11-02YJ01,2012BAD04B01-02)Key Science and Technology Program of Heilongjiang Province+2 种基金China(GA13B101)Heilongjiang Postdoctoral Sustentation Fund(LBH-Z10038)the Funds of Heilongjiang Academy of Agricultural Sciences for Distinguished Young Scholars(2014)~~