基金This research was funded by the grant fromthe National Natural Science Foundation of China (30499341 ,30270221) the Excellent YouthFoundationin Anhui Province (04043050)+2 种基金the Natural Science Foundation of Educational Committee of Anhui Province (2004sys003) the Foundation of Provincial Key Laboratory of Conservation and Utilizationfor I mportant Biological Resourcein Anhui the Special Foun-dation of Anhui Normal University