Partial pressure of CO2(pCO2) was investigated in the Changjiang(Yangtze River) Estuary,Hangzhou Bay and their adjacent areas during a cruise in August 2004,China.The data show that pCO2 in surface waters of the studi...Partial pressure of CO2(pCO2) was investigated in the Changjiang(Yangtze River) Estuary,Hangzhou Bay and their adjacent areas during a cruise in August 2004,China.The data show that pCO2 in surface waters of the studied area was higher than that in the atmosphere with only exception of a patch east of Zhoushan Archipelago.The pCO2 varied from 168 to 2 264 μatm,which fell in the low range compared with those of other estuaries in the world.The calculated sea-air CO2 fluxes decreased offshore and varied from -10.0 to 88.1 mmol m-2 d-1 in average of 24.4 ± 16.5 mmol m-2 d-1.Although the area studied was estimated only 2 × 104 km2,it emitted(5.9 ± 4.0) × 103 tons of carbon to the atmosphere every day.The estuaries and their plumes must be further studied for better understanding the role of coastal seas playing in the global oceanic carbon cycle.展开更多
The effect of hydrostatic pressure on the vibration dispersion characteristics of fluid-shell coupled structures was studied.Both fluid-loaded cylindrical shells and fluid-filled cylindrical shells were considered.Num...The effect of hydrostatic pressure on the vibration dispersion characteristics of fluid-shell coupled structures was studied.Both fluid-loaded cylindrical shells and fluid-filled cylindrical shells were considered.Numerical analysis was applied to solve the dispersion equations for shells filled with or loaded with fluid at various hydrostatic pressures.The results for external pressure showed that non-dimensional axial wave numbers are nearly independent when the pressure is below the critical level.The influence of internal pressure on wave numbers was found significant for the real branch s=1 and the complex branches of dispersion curves.The presence of internal pressure increased the cut on frequencies for the branch s=1 for high order wave modes.展开更多
Advantages of the detached phenomena have influenced researchers to modify the conventional methods to promote it on the earth. Since 1994, the vertical directional solidification (VDS) technique has been employed f...Advantages of the detached phenomena have influenced researchers to modify the conventional methods to promote it on the earth. Since 1994, the vertical directional solidification (VDS) technique has been employed for the growth of bulk crystals, without the seed, without contact to the ampoule wall, without coating and without external pressure. An automated furnace was designed and fabricated for the controlled temperature gradients, growth conditions and parameters. The typical ingots growths of GaSb have shown the gap of 20 μm-145 μm and mobility μn = 1125 cm^2/V.sec at 300 K. Mobility is highest and five times larger than the attached growths. Dislocation density is the order of 104/cm2 in the conical region, decreases in the direction of growth, and in many crystals reached less than 103/cm2. The spontaneous gap formation due to the meniscus depends on the pressure differences and thermal state. GaSb grown ingots have shown progress in the properties of crystal grown ever, and attributed to reduce thermal stress without contact to the ampoule wall.展开更多
The friction coefficient between tube and die in guide zone of tube hydroforming was obtained. In hydroforming, the tube is expanded by an internal pressure against the tool wall. By pushing the tube through tool, a f...The friction coefficient between tube and die in guide zone of tube hydroforming was obtained. In hydroforming, the tube is expanded by an internal pressure against the tool wall. By pushing the tube through tool, a friction force at the contact surface between the tube and the tool occurs. In guiding zone, the friction coefficients between tube and die can be estimated from the measured axial feeding forces. In expansion zone, the friction coefficients between tube and die can be evaluated from the measured geometries of expanded tubes and FE analysis.展开更多
A three-dimensional finite element model of the lumbar motion segment was used to predict the stress distribution in lumbar spine with posterior element resection. It was shown that the stress level in all parts of th...A three-dimensional finite element model of the lumbar motion segment was used to predict the stress distribution in lumbar spine with posterior element resection. It was shown that the stress level in all parts of the lumbar spine was elevated although the stress distribution remained unchanged. The authors concluded that the posterior element resection in lumbar surgery should be avoided as much as possible.展开更多
Thrust bearing is a key component of the propulsion system of a ship. It transfers the propulsive forces from the propeller to the ship's hull, allowing the propeller to push the ship ahead. The performance of a thru...Thrust bearing is a key component of the propulsion system of a ship. It transfers the propulsive forces from the propeller to the ship's hull, allowing the propeller to push the ship ahead. The performance of a thrust bearing pad is critical. When the thrust bearing becomes damaged, it can cause the ship to lose power and can also affect its operational safety. For this paper, the distribution of the pressure field of a thrust pad was calculated with numerical method, applying Reynolds equation. Thrust bearing properties for loads were analyzed, given variations in outlet thickness of the pad and variations between the load and the slope of the pad. It was noticed that the distribution of pressure was uneven. As a result, increases of both the outlet thickness and the slope coefficient of the pad were able to improve load beating capability.展开更多
For evaluate the aerodynamic character of the turbine cascades which have the aft-loaded profile, the experimental investigation was carried out on the low speed annular wind tunnel. And the detailed measurements of t...For evaluate the aerodynamic character of the turbine cascades which have the aft-loaded profile, the experimental investigation was carried out on the low speed annular wind tunnel. And the detailed measurements of the aerodynamic parameters were made from upstream to downstream of the two type turbine cascades, the one is the conventional straight blades cascade, the other is the curved blades cascades. The static pressure distributions on the endwall and the blade surface were also carried out. The influence of the aft-loaded profile and the curved blade on the development of loss and the pressure distribution was discussed, and analyses the different flow phenomena and mechanism in two type turbine cascades.展开更多
AIM: To investigate the flow and mixing at the duodenal stump after gastric resection, a computer simulation was implemented. METHODS: Using the finite element method, two different Billroth fl procedure cases (A a...AIM: To investigate the flow and mixing at the duodenal stump after gastric resection, a computer simulation was implemented. METHODS: Using the finite element method, two different Billroth fl procedure cases (A and B) were modeled. Case A was defined with a shorter and almost straight duodenal section, while case B has a much longer and curved duodenal section. Velocity, pressure and food concentration distribution were determined and the numerical results were compared with experimental observations. RESULTS: The pressure distribution obtained by numerical simulation was in the range of the recorded experimental results. Case A had a more favorable pressure distribution in comparison with case B. However, case B had better performance in terms of food transport because of more continual food distribution, as well as better emptying of the duodena section. CONCLUSION: This study offers insight into the transport process within the duodenal stump section after surgical intervention, which can be useful for future patient-specific predictions of a surgical outcome.展开更多
In this paper, by using adequate stress-strain relationship, mesh elements, boundary conditions and loading conditions, the finite element ANSYS analysis on the behavior of circular tubes subjected to symmetrical cycl...In this paper, by using adequate stress-strain relationship, mesh elements, boundary conditions and loading conditions, the finite element ANSYS analysis on the behavior of circular tubes subjected to symmetrical cyclic bending with or without external pressure is discussed. The behavior includes the moment-curvature and ovalization-curvature relationships. In addition, the calculated ovalizations at two different sections, middle and right cross-sections, are also included. Experimental data for 6061-T6 aluminum alloy tubes subjected to cyclic bending with or without external pressure were compared with the ANSYS analysis. It has been shown that the analysis of the elastoplatic moment-curvature relationship and the symmetrical, ratcheting and increasing ovalization-curvature relationship is in good agreement with the experimental data.展开更多
Based on the screening analysis for the first time in Montenegro, water samples for the presence of emergent wereanalyzed. Results from the locality Vukovci, the lower course of the river Morale, have shown the presen...Based on the screening analysis for the first time in Montenegro, water samples for the presence of emergent wereanalyzed. Results from the locality Vukovci, the lower course of the river Morale, have shown the presence of chemical components that indicate the presence of emergent in the water. Thus identification component is associated with the presence of fiziological groups of microorganisms at the site, which with its functional and significant role in many ways reduce environmental stress.展开更多
The paper inverts the focal mechanism solutions of the Yutian M_S7. 3 main shock,foreshocks and M_S≥3. 5 aftershocks by using the CAP method,based on the broadband waveforms recorded by the Xinjiang and Tibet Digital...The paper inverts the focal mechanism solutions of the Yutian M_S7. 3 main shock,foreshocks and M_S≥3. 5 aftershocks by using the CAP method,based on the broadband waveforms recorded by the Xinjiang and Tibet Digital Seismic Networks. The results show that the M_S7. 3 strong earthquake is of strike-slip type with a normal faulting component,and combined with the analysis of focal structure and the aftershock distribution,the nodal plane I with strike 241°,dip 90° and rake- 22° is considered to be the seismogenic fault plane of the main shock. The direction of P-axis for the main shock is 194°,close to the near NS direction of the principal stress P-axis of historical strong earthquakes in this region. The focal mechanism solution of the M_S5. 4 foreshock has a good consistency with that of the main shock. Among the 18 aftershocks,10 are of strike-slip type,6 are of normal faulting type and 2 are of thrust type. 70% of the aftershocks in the sequence have a focal mechanism with P-axis in the near-NS direction. The focal depths of this M_S7. 3 earthquake sequences are distributed in the range of 5km- 28 km,with the majority in the depth range of 15km- 20 km,slightly deeper than the depth of 10 km of the main shock as calculated.展开更多
Techniques of rotating-disk and catalyst were used in investigating the kinetics of dolomite dissolution in flowing CO2-H2O system. Experiments run in the solutions equilibrated with various CO2 partial pressures (PCO...Techniques of rotating-disk and catalyst were used in investigating the kinetics of dolomite dissolution in flowing CO2-H2O system. Experiments run in the solutions equilibrated with various CO2 partial pressures (PCO2) from 30 to 100000 Pa. It shows that dissolution rates ofdolomite are related with rotating speeds at conditions far from equilibrium. This was explained by modified diffusion boundary layer (DBL) model. In addition, the dissolution rates increase after addition of carbonic anhydrase (CA) to solutions, where the CA catalyzes CO2 conversion. However, great differences occur among various CO2 partial pressures. The experimental observations give a conclusion that the modified DBL model enables one to predict dissolution rates and their behaviour at various PCO2 with satisfactory precision at least far from equilibrium.展开更多
The velocity field of generalized second order fluid with fractional anomalous diiusion caused by a plate moving impulsively in its own plane is investigated and the anomalous diffusion problems of the stress field an...The velocity field of generalized second order fluid with fractional anomalous diiusion caused by a plate moving impulsively in its own plane is investigated and the anomalous diffusion problems of the stress field and vortex sheet caused by this process are studied. Many previous and classical results can be considered as particular cases of this paper, such as the solutions of the fractional diffusion equations obtained by Wyss; the classical Rayleigh’s time-space similarity solution; the relationship between stress field and velocity field obtained by Bagley and co-worker and Podlubny’s results on the fractional motion equation of a plate. In addition, a lot of significant results also are obtained. For example, the necessary condition for causing the vortex sheet is that the time fractional diffusion index β must be greater than that of generalized second order fluid α; the establiihment of the vorticity distribution function depends on the time history of the velocity profile at a given point, and the time history can be described by the fractional calculus.展开更多
Objective: To present our experience in using decompressive craniectomy (DC) among severe trau- matic brain injury (TBI) patients during operation and to discuss its indication. Methods: From October 2008 to Ma...Objective: To present our experience in using decompressive craniectomy (DC) among severe trau- matic brain injury (TBI) patients during operation and to discuss its indication. Methods: From October 2008 to May 2009, 41 patients aged between 18 and 75 years with severe TBI were in- cluded in this study. They underwent DC or non-DC (NDC) according to their intraoperative findings. Postoperative intracranial pressure (ICP), complications, requiring second operation or not and outcomes were observed. Results: Fifteen patients underwent DC and 26 pa- tients did not. The average postoperative ICP of each pa- tient was lower than 20 mm Hg. For patients received DC, 2 had seizures after operation and 1 developed cerebrocele in the follow-up period; only matic seizures, but none of 1 NDC patient had post-trau- them had delayed haematoma, cerebrospinal fluid fistula, cerebrocele or infections. At the end of follow-up, 10 patients died, 6 had the GOS of 2, 2 of 3, 9 of 4 and 14of 5. Conclusions: DC is necessary to manage fulminant intracranial hypertension or intraoperative brain swelling. If there was not brain swelling after removal of the haematoma and necrotized neural tissues, it is safe to replace skull flap. The intraoperative finding is an important factor to decide whether to perform DC or not.展开更多
As complex traits evolve, each component of the trait may be under different selection pressures and could respond independently to distinct evolutionary forces. We used comparative methods to examine patterns of evol...As complex traits evolve, each component of the trait may be under different selection pressures and could respond independently to distinct evolutionary forces. We used comparative methods to examine patterns of evolution in multiple components of a complex courtship signal in darters, specifically addressing the question of how nuptial coloration evolves across different areas of the body. Using spectral reflectance, we defined 4 broad color classes present on the body and fins of 17 species of freshwater fishes (genus Etheostoma) and quantified differences in hue within each color class. Ancestral state reconstruction suggests that most color traits were expressed in the most recent common ancestor of sampled species and that differences among species are mostly due to losses in coloration. The evolutionary lability of coloration varied across body regions; we found sig- nificant phylogenetic signal for orange color on the body but not for most colors on fins. Finally, patterns of color evolution and hue Of the colors were correlated among the two dorsal fins and between the anterior dorsal and anal fins, but not between any of the fins and the body. The observed patterns support the hypothesis that different components of complex signals may be subject to distinct evolutionary pressures, and suggests that the combination of behavioral displays and morphology in communication may have a strong influence on patterns of signal evolution .展开更多
基金Supported by the National Basic Research Program of China (973 Program, No. 2005CB407305)the open foundation of the Key Laboratory of Marine Sedimentology & Environmental Geology, SOA (No. MASEG200606)+1 种基金the Qingdao Special Program for Leading Scientists (No. 05-2-JC-90)the 100-Talent Project of the Chinese Academy of Sciences
文摘Partial pressure of CO2(pCO2) was investigated in the Changjiang(Yangtze River) Estuary,Hangzhou Bay and their adjacent areas during a cruise in August 2004,China.The data show that pCO2 in surface waters of the studied area was higher than that in the atmosphere with only exception of a patch east of Zhoushan Archipelago.The pCO2 varied from 168 to 2 264 μatm,which fell in the low range compared with those of other estuaries in the world.The calculated sea-air CO2 fluxes decreased offshore and varied from -10.0 to 88.1 mmol m-2 d-1 in average of 24.4 ± 16.5 mmol m-2 d-1.Although the area studied was estimated only 2 × 104 km2,it emitted(5.9 ± 4.0) × 103 tons of carbon to the atmosphere every day.The estuaries and their plumes must be further studied for better understanding the role of coastal seas playing in the global oceanic carbon cycle.
文摘The effect of hydrostatic pressure on the vibration dispersion characteristics of fluid-shell coupled structures was studied.Both fluid-loaded cylindrical shells and fluid-filled cylindrical shells were considered.Numerical analysis was applied to solve the dispersion equations for shells filled with or loaded with fluid at various hydrostatic pressures.The results for external pressure showed that non-dimensional axial wave numbers are nearly independent when the pressure is below the critical level.The influence of internal pressure on wave numbers was found significant for the real branch s=1 and the complex branches of dispersion curves.The presence of internal pressure increased the cut on frequencies for the branch s=1 for high order wave modes.
文摘Advantages of the detached phenomena have influenced researchers to modify the conventional methods to promote it on the earth. Since 1994, the vertical directional solidification (VDS) technique has been employed for the growth of bulk crystals, without the seed, without contact to the ampoule wall, without coating and without external pressure. An automated furnace was designed and fabricated for the controlled temperature gradients, growth conditions and parameters. The typical ingots growths of GaSb have shown the gap of 20 μm-145 μm and mobility μn = 1125 cm^2/V.sec at 300 K. Mobility is highest and five times larger than the attached growths. Dislocation density is the order of 104/cm2 in the conical region, decreases in the direction of growth, and in many crystals reached less than 103/cm2. The spontaneous gap formation due to the meniscus depends on the pressure differences and thermal state. GaSb grown ingots have shown progress in the properties of crystal grown ever, and attributed to reduce thermal stress without contact to the ampoule wall.
基金supported by grants-in-aid for the National Core Research Center Program from MEST/KOSEF
文摘The friction coefficient between tube and die in guide zone of tube hydroforming was obtained. In hydroforming, the tube is expanded by an internal pressure against the tool wall. By pushing the tube through tool, a friction force at the contact surface between the tube and the tool occurs. In guiding zone, the friction coefficients between tube and die can be estimated from the measured axial feeding forces. In expansion zone, the friction coefficients between tube and die can be evaluated from the measured geometries of expanded tubes and FE analysis.
文摘A three-dimensional finite element model of the lumbar motion segment was used to predict the stress distribution in lumbar spine with posterior element resection. It was shown that the stress level in all parts of the lumbar spine was elevated although the stress distribution remained unchanged. The authors concluded that the posterior element resection in lumbar surgery should be avoided as much as possible.
基金Supported by the Natural Science Foundation of China under Grant No.50675162the Program of Introducing Talents of Discipline to Universities under Grant No.B08031the Key Project of Hubei Province Science & Technology Fund under Grant No.2008CAD027
文摘Thrust bearing is a key component of the propulsion system of a ship. It transfers the propulsive forces from the propeller to the ship's hull, allowing the propeller to push the ship ahead. The performance of a thrust bearing pad is critical. When the thrust bearing becomes damaged, it can cause the ship to lose power and can also affect its operational safety. For this paper, the distribution of the pressure field of a thrust pad was calculated with numerical method, applying Reynolds equation. Thrust bearing properties for loads were analyzed, given variations in outlet thickness of the pad and variations between the load and the slope of the pad. It was noticed that the distribution of pressure was uneven. As a result, increases of both the outlet thickness and the slope coefficient of the pad were able to improve load beating capability.
文摘For evaluate the aerodynamic character of the turbine cascades which have the aft-loaded profile, the experimental investigation was carried out on the low speed annular wind tunnel. And the detailed measurements of the aerodynamic parameters were made from upstream to downstream of the two type turbine cascades, the one is the conventional straight blades cascade, the other is the curved blades cascades. The static pressure distributions on the endwall and the blade surface were also carried out. The influence of the aft-loaded profile and the curved blade on the development of loss and the pressure distribution was discussed, and analyses the different flow phenomena and mechanism in two type turbine cascades.
基金Supported by The Ministry of Science of Serbia with the grants OI144028 and TR12007
文摘AIM: To investigate the flow and mixing at the duodenal stump after gastric resection, a computer simulation was implemented. METHODS: Using the finite element method, two different Billroth fl procedure cases (A and B) were modeled. Case A was defined with a shorter and almost straight duodenal section, while case B has a much longer and curved duodenal section. Velocity, pressure and food concentration distribution were determined and the numerical results were compared with experimental observations. RESULTS: The pressure distribution obtained by numerical simulation was in the range of the recorded experimental results. Case A had a more favorable pressure distribution in comparison with case B. However, case B had better performance in terms of food transport because of more continual food distribution, as well as better emptying of the duodena section. CONCLUSION: This study offers insight into the transport process within the duodenal stump section after surgical intervention, which can be useful for future patient-specific predictions of a surgical outcome.
文摘In this paper, by using adequate stress-strain relationship, mesh elements, boundary conditions and loading conditions, the finite element ANSYS analysis on the behavior of circular tubes subjected to symmetrical cyclic bending with or without external pressure is discussed. The behavior includes the moment-curvature and ovalization-curvature relationships. In addition, the calculated ovalizations at two different sections, middle and right cross-sections, are also included. Experimental data for 6061-T6 aluminum alloy tubes subjected to cyclic bending with or without external pressure were compared with the ANSYS analysis. It has been shown that the analysis of the elastoplatic moment-curvature relationship and the symmetrical, ratcheting and increasing ovalization-curvature relationship is in good agreement with the experimental data.
文摘Based on the screening analysis for the first time in Montenegro, water samples for the presence of emergent wereanalyzed. Results from the locality Vukovci, the lower course of the river Morale, have shown the presence of chemical components that indicate the presence of emergent in the water. Thus identification component is associated with the presence of fiziological groups of microorganisms at the site, which with its functional and significant role in many ways reduce environmental stress.
基金funded jointly by Foundation of Earthquake Administration of Xinjiang Uygur Autonomous Region(Grant No.201401)the Contract for Annual Earthquake Situation Tracking Task of 2014,CEA(2014020106)
文摘The paper inverts the focal mechanism solutions of the Yutian M_S7. 3 main shock,foreshocks and M_S≥3. 5 aftershocks by using the CAP method,based on the broadband waveforms recorded by the Xinjiang and Tibet Digital Seismic Networks. The results show that the M_S7. 3 strong earthquake is of strike-slip type with a normal faulting component,and combined with the analysis of focal structure and the aftershock distribution,the nodal plane I with strike 241°,dip 90° and rake- 22° is considered to be the seismogenic fault plane of the main shock. The direction of P-axis for the main shock is 194°,close to the near NS direction of the principal stress P-axis of historical strong earthquakes in this region. The focal mechanism solution of the M_S5. 4 foreshock has a good consistency with that of the main shock. Among the 18 aftershocks,10 are of strike-slip type,6 are of normal faulting type and 2 are of thrust type. 70% of the aftershocks in the sequence have a focal mechanism with P-axis in the near-NS direction. The focal depths of this M_S7. 3 earthquake sequences are distributed in the range of 5km- 28 km,with the majority in the depth range of 15km- 20 km,slightly deeper than the depth of 10 km of the main shock as calculated.
基金the National Natural Science Foundation of China (Grant No.40073026), the Ministry of Land and Resources of China (Grant No. 9806), the Ministry of Science and Technology of China (Special Research Project for Social Commonweal) (Grant No. 164), the Na
文摘Techniques of rotating-disk and catalyst were used in investigating the kinetics of dolomite dissolution in flowing CO2-H2O system. Experiments run in the solutions equilibrated with various CO2 partial pressures (PCO2) from 30 to 100000 Pa. It shows that dissolution rates ofdolomite are related with rotating speeds at conditions far from equilibrium. This was explained by modified diffusion boundary layer (DBL) model. In addition, the dissolution rates increase after addition of carbonic anhydrase (CA) to solutions, where the CA catalyzes CO2 conversion. However, great differences occur among various CO2 partial pressures. The experimental observations give a conclusion that the modified DBL model enables one to predict dissolution rates and their behaviour at various PCO2 with satisfactory precision at least far from equilibrium.
基金the Doctoral Program Foundation of the Education Ministry of China the National Natural Science Foundation of China (Grant No. 10002003) Foundation for University Key Teacher by the Ministry of Education of China.
文摘The velocity field of generalized second order fluid with fractional anomalous diiusion caused by a plate moving impulsively in its own plane is investigated and the anomalous diffusion problems of the stress field and vortex sheet caused by this process are studied. Many previous and classical results can be considered as particular cases of this paper, such as the solutions of the fractional diffusion equations obtained by Wyss; the classical Rayleigh’s time-space similarity solution; the relationship between stress field and velocity field obtained by Bagley and co-worker and Podlubny’s results on the fractional motion equation of a plate. In addition, a lot of significant results also are obtained. For example, the necessary condition for causing the vortex sheet is that the time fractional diffusion index β must be greater than that of generalized second order fluid α; the establiihment of the vorticity distribution function depends on the time history of the velocity profile at a given point, and the time history can be described by the fractional calculus.
文摘Objective: To present our experience in using decompressive craniectomy (DC) among severe trau- matic brain injury (TBI) patients during operation and to discuss its indication. Methods: From October 2008 to May 2009, 41 patients aged between 18 and 75 years with severe TBI were in- cluded in this study. They underwent DC or non-DC (NDC) according to their intraoperative findings. Postoperative intracranial pressure (ICP), complications, requiring second operation or not and outcomes were observed. Results: Fifteen patients underwent DC and 26 pa- tients did not. The average postoperative ICP of each pa- tient was lower than 20 mm Hg. For patients received DC, 2 had seizures after operation and 1 developed cerebrocele in the follow-up period; only matic seizures, but none of 1 NDC patient had post-trau- them had delayed haematoma, cerebrospinal fluid fistula, cerebrocele or infections. At the end of follow-up, 10 patients died, 6 had the GOS of 2, 2 of 3, 9 of 4 and 14of 5. Conclusions: DC is necessary to manage fulminant intracranial hypertension or intraoperative brain swelling. If there was not brain swelling after removal of the haematoma and necrotized neural tissues, it is safe to replace skull flap. The intraoperative finding is an important factor to decide whether to perform DC or not.
基金Acknowledgements We thank Michael Martin, Tory Williams, and Alex Nahm for assistance in collecting fishes and the Mendelson lab for assistance in fish maintenance. We thank Chioma Ihekweazu for assistance with spectral analysis. We also thank Karen Carleton for use of the software to calculate spectral location. Tom Cronin, Kevin Omland, Megan Porter, Kate Feller, Nick Friedman and Brian Dalton provided helpful discussions throughout the course of this work. We thank two anonymous reviewers and the editor for their helpful comments on previous versions of this manuscript. We especially thank Eileen Hebets for the invitation to submit to this special issue. This work was funded through a National Science Foundation grant to TCM (#DEB 0718987). Additional support for this work came from the NSF, NCEAS and NESCENT funded "Comparative Phylogenetics in R" workshop attended by JMG.
文摘As complex traits evolve, each component of the trait may be under different selection pressures and could respond independently to distinct evolutionary forces. We used comparative methods to examine patterns of evolution in multiple components of a complex courtship signal in darters, specifically addressing the question of how nuptial coloration evolves across different areas of the body. Using spectral reflectance, we defined 4 broad color classes present on the body and fins of 17 species of freshwater fishes (genus Etheostoma) and quantified differences in hue within each color class. Ancestral state reconstruction suggests that most color traits were expressed in the most recent common ancestor of sampled species and that differences among species are mostly due to losses in coloration. The evolutionary lability of coloration varied across body regions; we found sig- nificant phylogenetic signal for orange color on the body but not for most colors on fins. Finally, patterns of color evolution and hue Of the colors were correlated among the two dorsal fins and between the anterior dorsal and anal fins, but not between any of the fins and the body. The observed patterns support the hypothesis that different components of complex signals may be subject to distinct evolutionary pressures, and suggests that the combination of behavioral displays and morphology in communication may have a strong influence on patterns of signal evolution .