基金Acknowledgements We are deeply grateful to Leng Cong and Long Ping for collecting specimens. This work was supported by the National Natural Sciences Foundation of China (Grant No. 31172113 / 31272268 / 31601882), the Special Foundation of Ministry of Science and Technology of the People' s Republic of China (Grant No. 2014FY 110100) and the Natural Science Foundation of Guizhou Province (Grant No. J [2014] 2146).