In this paper, a new species of the genus Oplitis Berlese, 1884, namely Oplitis tielingensis sp. nov. from Longshou Mountain, Tieling City, Liaoning Province, China is described and figured. The new species is similar...In this paper, a new species of the genus Oplitis Berlese, 1884, namely Oplitis tielingensis sp. nov. from Longshou Mountain, Tieling City, Liaoning Province, China is described and figured. The new species is similar to Oplitis yuxini Ma, 2001. The type specimens are deposited in the College of Protection, Shenyang Agricultural University.展开更多
The new genus Graminothrips and two new species G. cyperi and G. longisetosus are described from China.In the course of sorting out the grass-living thrips of South China, the authors found some interesting specimens....The new genus Graminothrips and two new species G. cyperi and G. longisetosus are described from China.In the course of sorting out the grass-living thrips of South China, the authors found some interesting specimens. They belong to a new genus, Graminothrips, and two new species as described below. All types are preserved in the Insect Collection of South China Agricultural University.展开更多
A new genus, Thelazacallis Zhang (Homoptera: Drepanosiphidae) and a new species, Thelazacallis ranunculicola Qiao et Zhang from China is described with 9 original figures in the present paper. All type specimens a...A new genus, Thelazacallis Zhang (Homoptera: Drepanosiphidae) and a new species, Thelazacallis ranunculicola Qiao et Zhang from China is described with 9 original figures in the present paper. All type specimens are deposited in the Institute of Zoology, Chinese Academy of Sciences.展开更多
A new leaf beetle genus Yulongedon gen. nov., and two new species, Y. formosus sp. nov. and Y. jambhalai sp. nov. from China, are described and illustrated here.
In the present paper, a new species, Placidellus conjugatus, sp. nov. is described from Fujian, China and its types are deposited in the Entomological Museum, Northwest SciTech University of Agriculture and Forestry (...In the present paper, a new species, Placidellus conjugatus, sp. nov. is described from Fujian, China and its types are deposited in the Entomological Museum, Northwest SciTech University of Agriculture and Forestry (NSUAF). One related new genus, Paraplacidellus, gen. nov. with a new species Paraplacidellus maculatus, sp. nov. is described from Sandakan, Borneo, and the holotype of the new species is deposited in the National Museum of Natural History (NMNH).展开更多
Spinarge, gen.nov.with two new species, Spinarge sichuanensis, sp.nov.and S. liui, sp.nov. are described. The new genus is placed in subfamily Athermantinae. A key to the two new species is provided.
The paper reports a new genus of Arcypteridae i e. Suacris gen.nov.and a new species Suacris siyangensis sp.nov.The new genus is similar to Omocestus I Bol ,but differs from the latter in:① Lateral car...The paper reports a new genus of Arcypteridae i e. Suacris gen.nov.and a new species Suacris siyangensis sp.nov.The new genus is similar to Omocestus I Bol ,but differs from the latter in:① Lateral carinae slightly curved,widest 1 6-1 8 times greater than narrowest.② Cubital area wider than or equal to medial area.③Tympanum organ distinct,♂ widely oval,♀ crevice like.The new genus is also similar to Stenobothroides Xu et Zheng,but differs from the later in:①Lateral carinae obvious and curve.②Tergum of terminal abdomere without furcula.③Male subgenital plate long conica.Type specimens are deposited in Northwest Plateau Institute of Biology,the Chinese Academy of Sciences.展开更多
A new genus and two new species from the family Tenebrionidae of the order Coleoptera of China are reported in this paper,namely Neoblaps gen.nov.,N.huizensis sp.nov.,Dila bomiana sp.nov.The type specimens are preserv...A new genus and two new species from the family Tenebrionidae of the order Coleoptera of China are reported in this paper,namely Neoblaps gen.nov.,N.huizensis sp.nov.,Dila bomiana sp.nov.The type specimens are preserved in the Museum of Hebei University.展开更多
A new Oriental dictyopharid genus and species, Dictyotenguna choui gen. n., sp. n., is described and illustrated from China. A brief review of the family Dictyopharidae from the Oriental and eastern Palaearctic Region...A new Oriental dictyopharid genus and species, Dictyotenguna choui gen. n., sp. n., is described and illustrated from China. A brief review of the family Dictyopharidae from the Oriental and eastern Palaearctic Regions is provided.展开更多
The new genus Parathrausta gen.n. in the subfamily Spilomelinae (Lepidoptera, Crambidae) is erected based on a series of specimens collected in the Afrotropical region of Saudi Arabia. The new genus is monotypical, wi...The new genus Parathrausta gen.n. in the subfamily Spilomelinae (Lepidoptera, Crambidae) is erected based on a series of specimens collected in the Afrotropical region of Saudi Arabia. The new genus is monotypical, with Parathrausta internervalis sp.n. its type species. The adults of the type species, the male genitalia, the female genitalia and the tympanal organs are described and figured. The new genus is recognized as related by characters in the male and female genitalia with genera in the tribes Steniini Guine, 1854 and Nomophilini Kuznetzov & Stekolnikov, 1979, namely with Bradina Lederer, 1863, Perisyntrocha Meyrick, 1894, Diathrausta Lederer, 1863, Diasemiopsis Munroe, 1957, Diasemiodes Munroe, 1957 and with the genera in the Duponchelia Zeller, 1847 group comprising Duponchelia Zeller, 1847, Penestola Mschler, 1890, Tatobotys Butler, 1880 and Hymenoptychis Zeller, 1852. The character states differentiating the new genus from the comparative genera are discussed. Potential autapomorphies of the new genus are listed. The tribal assignment of the new genus to the Nomophilini Kuznetzov & Stekolnikov, 1979 based on morphological characters is discussed. The support of the tribal and cladistic assignment of the new genus as a sister clade to Diathrausta Lederer, 1863 based phylogenetic analyses (NJ, MP and ML) is shown. An unusually strong genetic divergence of the genus in the COI segment of the mt-DNA based on the Tamura-Nei distance measure from the morphologically related genera is recognized and addressed.展开更多
A new genus and species of threefrog is described from Medog, southeastern Tibet, China based on morphological and phylogenetic data. The new genus can be distinguished from other treefrog genera by the following comb...A new genus and species of threefrog is described from Medog, southeastern Tibet, China based on morphological and phylogenetic data. The new genus can be distinguished from other treefrog genera by the following combination of characters： （1） body size moderate, 45.0 mm in male; （2） snout rounded; （3） canthus rostralis obtuse and raised prominently, forming a ridge from nostril to anterior corner of eyes; （4） web rudimentary on fingers; （5） web moderately developed on toes; （6） phalange ＂Y＂ shaped, visible from dorsal side of fingers and toes; （7） skin of dorsal surfaces relatively smooth, scatted with small tubercles; （8） iris with a pale yellow, ＂X＂ shaped pattern of pigmentation.展开更多
Two new water mite （Acari： Hygrobatidae） species are described from Sichuan Province, China, Atractides （Atractides） yaaniensis sp. nov. and Atractides （Atractides） yaaniensisoides sp. nov.. The materials were ...Two new water mite （Acari： Hygrobatidae） species are described from Sichuan Province, China, Atractides （Atractides） yaaniensis sp. nov. and Atractides （Atractides） yaaniensisoides sp. nov.. The materials were collected in 2011 during a water mite survey by Zeyu WANG from a river in Sichuan.展开更多
This study describes and illustrates four new species Laticanna nanhaiensis Xu, Huang and Wang, gen. nov. and sp. nov., Protiaropsis gemmifera Xu, Huang and Du, sp. nov., Merga nanhaiensis Xu, Huang and Guo, sp. nov. ...This study describes and illustrates four new species Laticanna nanhaiensis Xu, Huang and Wang, gen. nov. and sp. nov., Protiaropsis gemmifera Xu, Huang and Du, sp. nov., Merga nanhaiensis Xu, Huang and Guo, sp. nov. and Rhabdoon apiciloculus Xu, Huang and Du, sp. nov. based on specimens from the south-central South China Sea. Additionally, keys to known genera of family Bythotiaridae and species of genera Protiaropsis, Merga, and Rhabdoon are described. All type specimens are deposited in the South China Sea Fisheries Research Institute, Chinese Academy of Fishery Science.展开更多
-A new genus and a new species have been established on the basis of three specimens collected from Fujian's coastal zone in China. This new genus is named Laevispinereis. Its brief characters are as follows: oral...-A new genus and a new species have been established on the basis of three specimens collected from Fujian's coastal zone in China. This new genus is named Laevispinereis. Its brief characters are as follows: oral ring of proboscis has papillae, maxillary ring has conical paragnaths; the peristomium greatly expands into a collar; notosetaes are homogomph spinigers, neurosetaes are homogomph spinigers and homogomph falcigers; tip of spinigers setae with minute serratures and smooth on side. The type of genus is Laevispinereis fujianensis sp. nov., it has conical paragnaths on maxillary ring of proboscis: I has Ⅰ; Ⅱ has 6-7: Ⅲ has 12;Ⅳ has 6-7.展开更多
The genus Beipiaoserphus gen. nov. is established and referred to Mesoserphidae with Beipiaoserphus elegans sp. nov. as its type species. The type specimens were collected from the lower part of the Upper Jura...The genus Beipiaoserphus gen. nov. is established and referred to Mesoserphidae with Beipiaoserphus elegans sp. nov. as its type species. The type specimens were collected from the lower part of the Upper Jurassic Yixian Formation at Huangbanjigou Village near Shangyuan Town, Beipiao, Liaoning Province, China and are housed at Nanjing Institute of Geology and Palaeontology, Academia Sinica, Nanjing.展开更多
A new genus of Proctotrupinae is established to receive Carinaserphus sinensis, sp. nov. from Mt. Funiu, Henan Province, China. The new genus is described and compared with related genera. The type specimen is deposit...A new genus of Proctotrupinae is established to receive Carinaserphus sinensis, sp. nov. from Mt. Funiu, Henan Province, China. The new genus is described and compared with related genera. The type specimen is deposited in the Hymenopteran Collection of Zhejiang University, China.展开更多
One new genus and four new species of eriophyoid mites from Hainan, China are described and illustrated in this paper： Dicoxiseta Wang, Tan ＆ Yang, gem nov., D. litsea Wang, Tan ＆ Yang, sp. nov. infesting Litsea mo...One new genus and four new species of eriophyoid mites from Hainan, China are described and illustrated in this paper： Dicoxiseta Wang, Tan ＆ Yang, gem nov., D. litsea Wang, Tan ＆ Yang, sp. nov. infesting Litsea monopetala （Roxb.） Pers. （Lauraceae）, Eriophyes casearius Wang, Yang ＆ Tan, sp. nov. infesting Casearia glomerata Roxb. （Flacourtiaceae）, Phyllocoptruta alstonia Wang, Gu ＆ Tan, sp. nov. infesting Alstonia scholaris （L.） R. Br. （Apocynaceae） and Aculus paederius Wang, Gu ＆ Tan, sp. nov. infesting Paederia scandens （Lour.） Merr. （Rubiaceae）. Tegolophus artocarpi Keifer, 1977 is newly recorded from China. E. casearius Wang, Yang ＆ Tan, sp. nov. distorts severely new foliage of its host plant. Three species, P. alstonia Wang, Gu ＆ Tan, sp. nov., A. paederius Wang, Gu ＆ Tan, sp. nov. and T. artocarpi, cause rust on the undersurfaces of host plants, while D. litsea Wang, Tan ＆ Yang, sp. nov. is a vagrant on the undersurface of host plant leaves, without visible symptom.展开更多
Previous studies recognized three major lineages of the family Costaceae:a South American clade,an Asian clade and a Costus clade.However,the genus Hellenia within the Asian clade has been shown to be non-monophyletic...Previous studies recognized three major lineages of the family Costaceae:a South American clade,an Asian clade and a Costus clade.However,the genus Hellenia within the Asian clade has been shown to be non-monophyletic and its morphology has not been studied carefully.Therefore,the complete plastid genomes of Hellenia species were obtained and the monophyly of Hellenia was tested through four different datasets in this study.Plastid phylogenomic analyses of Costaceae revealed that Hellenia is strongly supported as paraphyletic.Two major clades are recovered,namely the Hellenia s.s.subclade and the Parahellenia subclade.Phylogenetic analyses based on an enlarged taxon sampling of the Asian clade using a two chloroplast markers dataset(trnK intron and trnL-F spacer)confirmed the paraphyly of Hellenia.Meanwhile,morphological analyses suggested that members of the Parahellenia subclade differ from the remaining Hellenia species in many characters including inflorescences,bracts,stigma,axillary buds,floral tubes and labellum.According to the present molecular and morphological evidence,the latter subclade is recognized as a new genus,Parahellenia.Two new species are described,four new combinations are made,and identification keys are also provided.展开更多
Parageotrupes incanus gen. et sp. nov. （Scarabaeoidea： Geotrupidae： Geotrupinae） is described and illustrated from the Yixian Formation of western Liaoning province, China.
文摘In this paper, a new species of the genus Oplitis Berlese, 1884, namely Oplitis tielingensis sp. nov. from Longshou Mountain, Tieling City, Liaoning Province, China is described and figured. The new species is similar to Oplitis yuxini Ma, 2001. The type specimens are deposited in the College of Protection, Shenyang Agricultural University.
文摘The new genus Graminothrips and two new species G. cyperi and G. longisetosus are described from China.In the course of sorting out the grass-living thrips of South China, the authors found some interesting specimens. They belong to a new genus, Graminothrips, and two new species as described below. All types are preserved in the Insect Collection of South China Agricultural University.
基金The work was supported by the National Natural Science Foundation of China!(No. 3970 0 0 1 5)
文摘A new genus, Thelazacallis Zhang (Homoptera: Drepanosiphidae) and a new species, Thelazacallis ranunculicola Qiao et Zhang from China is described with 9 original figures in the present paper. All type specimens are deposited in the Institute of Zoology, Chinese Academy of Sciences.
基金supported by a grant from the National Science Foundation of China to Xingke YANG(PI, 31010103913)Siqin GE(PI,30970390)+2 种基金grants from the Key Laboratory of Zoological Systematics and Evolution of the Chinese Academy of Sciences(O529YX5105)the Knowledge Innovation Program of Chinese Academy of Sciences(KSCX3-IOZ-1004)the Chinese National Science Fund for Fostering Talents in Basic Research(Special Subjects in Animal Taxonomy,NSFC-J0630964 /J0109,J0930004)
文摘A new leaf beetle genus Yulongedon gen. nov., and two new species, Y. formosus sp. nov. and Y. jambhalai sp. nov. from China, are described and illustrated here.
基金This study is supported by the National Natural Science Foundation of China( 39870 1 1 3)
文摘In the present paper, a new species, Placidellus conjugatus, sp. nov. is described from Fujian, China and its types are deposited in the Entomological Museum, Northwest SciTech University of Agriculture and Forestry (NSUAF). One related new genus, Paraplacidellus, gen. nov. with a new species Paraplacidellus maculatus, sp. nov. is described from Sandakan, Borneo, and the holotype of the new species is deposited in the National Museum of Natural History (NMNH).
文摘Spinarge, gen.nov.with two new species, Spinarge sichuanensis, sp.nov.and S. liui, sp.nov. are described. The new genus is placed in subfamily Athermantinae. A key to the two new species is provided.
文摘The paper reports a new genus of Arcypteridae i e. Suacris gen.nov.and a new species Suacris siyangensis sp.nov.The new genus is similar to Omocestus I Bol ,but differs from the latter in:① Lateral carinae slightly curved,widest 1 6-1 8 times greater than narrowest.② Cubital area wider than or equal to medial area.③Tympanum organ distinct,♂ widely oval,♀ crevice like.The new genus is also similar to Stenobothroides Xu et Zheng,but differs from the later in:①Lateral carinae obvious and curve.②Tergum of terminal abdomere without furcula.③Male subgenital plate long conica.Type specimens are deposited in Northwest Plateau Institute of Biology,the Chinese Academy of Sciences.
文摘A new genus and two new species from the family Tenebrionidae of the order Coleoptera of China are reported in this paper,namely Neoblaps gen.nov.,N.huizensis sp.nov.,Dila bomiana sp.nov.The type specimens are preserved in the Museum of Hebei University.
基金supported by the following sources:National Natural Science Foundation of China (31101657,30970400,31172128)Scientific Survey on the Middle- and Lower-reaches of Lancang (Mekong) River and Grand Shangri-La Area (2008FY110300)+1 种基金National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy,NSFC-J0630964/J0109)a grant (O529YX5105) from the Key Laboratory of Zoological Systematics and Evolution of the Chinese Academy of Sciences
文摘A new Oriental dictyopharid genus and species, Dictyotenguna choui gen. n., sp. n., is described and illustrated from China. A brief review of the family Dictyopharidae from the Oriental and eastern Palaearctic Regions is provided.
文摘The new genus Parathrausta gen.n. in the subfamily Spilomelinae (Lepidoptera, Crambidae) is erected based on a series of specimens collected in the Afrotropical region of Saudi Arabia. The new genus is monotypical, with Parathrausta internervalis sp.n. its type species. The adults of the type species, the male genitalia, the female genitalia and the tympanal organs are described and figured. The new genus is recognized as related by characters in the male and female genitalia with genera in the tribes Steniini Guine, 1854 and Nomophilini Kuznetzov & Stekolnikov, 1979, namely with Bradina Lederer, 1863, Perisyntrocha Meyrick, 1894, Diathrausta Lederer, 1863, Diasemiopsis Munroe, 1957, Diasemiodes Munroe, 1957 and with the genera in the Duponchelia Zeller, 1847 group comprising Duponchelia Zeller, 1847, Penestola Mschler, 1890, Tatobotys Butler, 1880 and Hymenoptychis Zeller, 1852. The character states differentiating the new genus from the comparative genera are discussed. Potential autapomorphies of the new genus are listed. The tribal assignment of the new genus to the Nomophilini Kuznetzov & Stekolnikov, 1979 based on morphological characters is discussed. The support of the tribal and cladistic assignment of the new genus as a sister clade to Diathrausta Lederer, 1863 based phylogenetic analyses (NJ, MP and ML) is shown. An unusually strong genetic divergence of the genus in the COI segment of the mt-DNA based on the Tamura-Nei distance measure from the morphologically related genera is recognized and addressed.
基金supported by the Ministry of Science and Technology of China(2014FY210200,2011FY120200)the Animal Branch of the Germplasm Bank of Wild Species of Chinese Academy of Sciences(the Large Research Infrastructure Funding)
文摘A new genus and species of threefrog is described from Medog, southeastern Tibet, China based on morphological and phylogenetic data. The new genus can be distinguished from other treefrog genera by the following combination of characters： （1） body size moderate, 45.0 mm in male; （2） snout rounded; （3） canthus rostralis obtuse and raised prominently, forming a ridge from nostril to anterior corner of eyes; （4） web rudimentary on fingers; （5） web moderately developed on toes; （6） phalange ＂Y＂ shaped, visible from dorsal side of fingers and toes; （7） skin of dorsal surfaces relatively smooth, scatted with small tubercles; （8） iris with a pale yellow, ＂X＂ shaped pattern of pigmentation.
基金funded by the National Natural Science Foundation of China (31071967)
文摘Two new water mite （Acari： Hygrobatidae） species are described from Sichuan Province, China, Atractides （Atractides） yaaniensis sp. nov. and Atractides （Atractides） yaaniensisoides sp. nov.. The materials were collected in 2011 during a water mite survey by Zeyu WANG from a river in Sichuan.
基金The National Natural Science Foundation of China under contract No.41406188the Grant of China Ministry of Science and Technology under contract No.2013BAD13B06+2 种基金the Grant of Guangdong Province Ministry of Science and Technology under contract No.2014A020217011the Ministry of Agriculture Major Funding Projects under contract No.NFZX2013the National Basic Research Program(973 Program)of China under contract No.2011CB403604
文摘This study describes and illustrates four new species Laticanna nanhaiensis Xu, Huang and Wang, gen. nov. and sp. nov., Protiaropsis gemmifera Xu, Huang and Du, sp. nov., Merga nanhaiensis Xu, Huang and Guo, sp. nov. and Rhabdoon apiciloculus Xu, Huang and Du, sp. nov. based on specimens from the south-central South China Sea. Additionally, keys to known genera of family Bythotiaridae and species of genera Protiaropsis, Merga, and Rhabdoon are described. All type specimens are deposited in the South China Sea Fisheries Research Institute, Chinese Academy of Fishery Science.
文摘-A new genus and a new species have been established on the basis of three specimens collected from Fujian's coastal zone in China. This new genus is named Laevispinereis. Its brief characters are as follows: oral ring of proboscis has papillae, maxillary ring has conical paragnaths; the peristomium greatly expands into a collar; notosetaes are homogomph spinigers, neurosetaes are homogomph spinigers and homogomph falcigers; tip of spinigers setae with minute serratures and smooth on side. The type of genus is Laevispinereis fujianensis sp. nov., it has conical paragnaths on maxillary ring of proboscis: I has Ⅰ; Ⅱ has 6-7: Ⅲ has 12;Ⅳ has 6-7.
基金Supported by the National Natural Science foundation of China (49832 0 2 0 NIGPAS990 50 2 ) and CMBRP(G2 0 0 0 0 7770 0 )
文摘The genus Beipiaoserphus gen. nov. is established and referred to Mesoserphidae with Beipiaoserphus elegans sp. nov. as its type species. The type specimens were collected from the lower part of the Upper Jurassic Yixian Formation at Huangbanjigou Village near Shangyuan Town, Beipiao, Liaoning Province, China and are housed at Nanjing Institute of Geology and Palaeontology, Academia Sinica, Nanjing.
文摘A new genus of Proctotrupinae is established to receive Carinaserphus sinensis, sp. nov. from Mt. Funiu, Henan Province, China. The new genus is described and compared with related genera. The type specimen is deposited in the Hymenopteran Collection of Zhejiang University, China.
基金supported by the Guangxi Special Invited Scientist Program in Agric-Environment and Agric-products Safety,the National Natural Science Foundation of China(31160431)the Key Project of Chinese Ministry of Education(211134)
文摘One new genus and four new species of eriophyoid mites from Hainan, China are described and illustrated in this paper： Dicoxiseta Wang, Tan ＆ Yang, gem nov., D. litsea Wang, Tan ＆ Yang, sp. nov. infesting Litsea monopetala （Roxb.） Pers. （Lauraceae）, Eriophyes casearius Wang, Yang ＆ Tan, sp. nov. infesting Casearia glomerata Roxb. （Flacourtiaceae）, Phyllocoptruta alstonia Wang, Gu ＆ Tan, sp. nov. infesting Alstonia scholaris （L.） R. Br. （Apocynaceae） and Aculus paederius Wang, Gu ＆ Tan, sp. nov. infesting Paederia scandens （Lour.） Merr. （Rubiaceae）. Tegolophus artocarpi Keifer, 1977 is newly recorded from China. E. casearius Wang, Yang ＆ Tan, sp. nov. distorts severely new foliage of its host plant. Three species, P. alstonia Wang, Gu ＆ Tan, sp. nov., A. paederius Wang, Gu ＆ Tan, sp. nov. and T. artocarpi, cause rust on the undersurfaces of host plants, while D. litsea Wang, Tan ＆ Yang, sp. nov. is a vagrant on the undersurface of host plant leaves, without visible symptom.
基金funded by the National Natural Science Foundation of China (Grant no.32070223)the Biological Resources Programme,Chinese Academy of Sciences (Grant no.KFJ-BRP-017-19)
文摘Previous studies recognized three major lineages of the family Costaceae:a South American clade,an Asian clade and a Costus clade.However,the genus Hellenia within the Asian clade has been shown to be non-monophyletic and its morphology has not been studied carefully.Therefore,the complete plastid genomes of Hellenia species were obtained and the monophyly of Hellenia was tested through four different datasets in this study.Plastid phylogenomic analyses of Costaceae revealed that Hellenia is strongly supported as paraphyletic.Two major clades are recovered,namely the Hellenia s.s.subclade and the Parahellenia subclade.Phylogenetic analyses based on an enlarged taxon sampling of the Asian clade using a two chloroplast markers dataset(trnK intron and trnL-F spacer)confirmed the paraphyly of Hellenia.Meanwhile,morphological analyses suggested that members of the Parahellenia subclade differ from the remaining Hellenia species in many characters including inflorescences,bracts,stigma,axillary buds,floral tubes and labellum.According to the present molecular and morphological evidence,the latter subclade is recognized as a new genus,Parahellenia.Two new species are described,four new combinations are made,and identification keys are also provided.
基金supported by grants from the National Natural Science Foundation of China(No.40872022)the Nature Science Foundation of Beijing(No.5082002)the Scientific Research Key Program KZ200910028005 and PHR20090509 Project of Beijing Municipal Commission of Education
文摘Parageotrupes incanus gen. et sp. nov. （Scarabaeoidea： Geotrupidae： Geotrupinae） is described and illustrated from the Yixian Formation of western Liaoning province, China.
