基金supported by the National Natural Science Foundation of China (61622406,61904015,11674310,61725505 and 11734016)the National Key Research and Development Program of China (2017YFA0207500)+2 种基金the Strategic Priority Research Program of Chinese Academy of Sciences (XDB30000000)the “The Pearl River Talent Recruitment Program”(2019ZT08X639)Beijing National Laboratory for Molecular Sciences (BNLMS201908)。