A new genus and two new species of fossil Fuziidae are described and illustrated: Parvifuzia gen. nov., P. marsa sp. nov. and P. brava sp. nov.. Both species were collected from the Middle Jurassic Jiulongshan, Forma...A new genus and two new species of fossil Fuziidae are described and illustrated: Parvifuzia gen. nov., P. marsa sp. nov. and P. brava sp. nov.. Both species were collected from the Middle Jurassic Jiulongshan, Formation of Daohugou, Inner Mongolia of China. In conjunction with Fuzia Vr(s)ansk(y),Liang et Ren, 2009, the present findings allow us to demonstrate that members of the family were morphologically diverse, with body length ranging from medium (17 mm) in Fuzia to small (9.7 mm) in the present genus. Wing venation of F. dadao is complicated, but simple in the present species, and apex of male's cerci are sharp and forceps-like in F. dadao, but strongly curved inward and round in the new genus.展开更多
A completely preserved cockroach Socialaperlucida gen. et sp. nov. (Blattida: Socialidae faro. nov.) is described from the Mesozoic (Albian Early Cretaceous) amber of Archingeay in France. It is categorized withi...A completely preserved cockroach Socialaperlucida gen. et sp. nov. (Blattida: Socialidae faro. nov.) is described from the Mesozoic (Albian Early Cretaceous) amber of Archingeay in France. It is categorized within the new family Socialidae, originating from the Mesozoic cockroach family Liberiblattinidae, and representing the sister group to the most basal known eutermite families. Numerous direct and indirect morphological evidence (such as a unique narrow body and pronotum with reduced coloration, legs without carination and with numerous sensillar pitts (forelegs) as adaptations to life in nests analogical to the living termitophillous species; enlarged head, long palps for communication, a general venation modified in a direction towards termites (subcosta and radial vein [R] approximated, R branches approximated, simplified, cross-veins reduced), and possibly also the development of the breaking forewing sutura for detaching wings after marriage flight), suggest its eusocial mode of life. Thus, the first eusociality originated within cockroaches, prior to giving birth to termites; termites and Cryptocercus are not directly related. The group of eusocial cockroaches makes termite morphotaxon polyphyletic, but also in this case, the erection of a new order for mastotermites (including Cratomastotermitidae) and/or the inclusion of eusocial, morphological cockroaches within termites appears counterproductive, thus an example of such a general exception to the taxonomical procedure is provided.展开更多
Vrsansky, (2000) considered there is decreasing variability in the forewings of cockroaches through time, based on a study of fossil and Recent species. This study tests his theory, based on a study of four Recent s...Vrsansky, (2000) considered there is decreasing variability in the forewings of cockroaches through time, based on a study of fossil and Recent species. This study tests his theory, based on a study of four Recent species of cockroaches: Blattella germanica (Blattellidae), Polyphaga aegyptiaca (Polyphagidae), Shelfordella lateralis (Blattidae) and Blaberus craniifer (Blaberidae). The new results, based on the coefficient of variation (CV) of the number of veins, indicate that Recent species can be more variable than fossil species and therefore do not support the theory. The results also show that at least 30 complete wings of the same species are required for a reliable CV value that is compa- rable between species.展开更多
Cariblattoides labandeirai sp.n. from the Eocene sediments of Green River in Colorado, USA bear only two plesiomorphies, but also several significant autapomorphies within the advanced and highly derived living cockro...Cariblattoides labandeirai sp.n. from the Eocene sediments of Green River in Colorado, USA bear only two plesiomorphies, but also several significant autapomorphies within the advanced and highly derived living cockroach genus. Thus, Cariblattoides with extant occurrence in the Caribbean and South America was historically common in the Nearctic, and represents important evidence for the occurrence of derived living genera of cockroaches -50 Ma ago. Generally, the vast majority of living genera were absent during the Palaeocene, thus the diversification of most living cockroach lineages near the Palaeocene/Eocene boundary must have been extremely rapid. Females of living C. suave, the type species, have identical (sophisticated) coloration ofpronotum, but the most related living taxa are C. piraiensis and C.fontesi from Brazil (supported by phylogenetical analysis).展开更多
基金supported by the National Natural Science Foundation of China (No. 40872022, 31071964)Nature Science Foundation of Beijing (No. 5082002)+1 种基金the Scientific Research Key Program (KZ200910028005)PHR Project of Beijing Municipal Commission of Education
文摘A new genus and two new species of fossil Fuziidae are described and illustrated: Parvifuzia gen. nov., P. marsa sp. nov. and P. brava sp. nov.. Both species were collected from the Middle Jurassic Jiulongshan, Formation of Daohugou, Inner Mongolia of China. In conjunction with Fuzia Vr(s)ansk(y),Liang et Ren, 2009, the present findings allow us to demonstrate that members of the family were morphologically diverse, with body length ranging from medium (17 mm) in Fuzia to small (9.7 mm) in the present genus. Wing venation of F. dadao is complicated, but simple in the present species, and apex of male's cerci are sharp and forceps-like in F. dadao, but strongly curved inward and round in the new genus.
基金supported by UNESCO-AMBA,UNESCO-IGCP 458,Schwarz stipend, VEGA 6002 and 2/0125/09,MVTS,and the Literarny Fond.a contribution to the CNRS ANR program"AMBRACE"(project no.BLAN07-1-184190)
文摘A completely preserved cockroach Socialaperlucida gen. et sp. nov. (Blattida: Socialidae faro. nov.) is described from the Mesozoic (Albian Early Cretaceous) amber of Archingeay in France. It is categorized within the new family Socialidae, originating from the Mesozoic cockroach family Liberiblattinidae, and representing the sister group to the most basal known eutermite families. Numerous direct and indirect morphological evidence (such as a unique narrow body and pronotum with reduced coloration, legs without carination and with numerous sensillar pitts (forelegs) as adaptations to life in nests analogical to the living termitophillous species; enlarged head, long palps for communication, a general venation modified in a direction towards termites (subcosta and radial vein [R] approximated, R branches approximated, simplified, cross-veins reduced), and possibly also the development of the breaking forewing sutura for detaching wings after marriage flight), suggest its eusocial mode of life. Thus, the first eusociality originated within cockroaches, prior to giving birth to termites; termites and Cryptocercus are not directly related. The group of eusocial cockroaches makes termite morphotaxon polyphyletic, but also in this case, the erection of a new order for mastotermites (including Cratomastotermitidae) and/or the inclusion of eusocial, morphological cockroaches within termites appears counterproductive, thus an example of such a general exception to the taxonomical procedure is provided.
文摘Vrsansky, (2000) considered there is decreasing variability in the forewings of cockroaches through time, based on a study of fossil and Recent species. This study tests his theory, based on a study of four Recent species of cockroaches: Blattella germanica (Blattellidae), Polyphaga aegyptiaca (Polyphagidae), Shelfordella lateralis (Blattidae) and Blaberus craniifer (Blaberidae). The new results, based on the coefficient of variation (CV) of the number of veins, indicate that Recent species can be more variable than fossil species and therefore do not support the theory. The results also show that at least 30 complete wings of the same species are required for a reliable CV value that is compa- rable between species.
文摘Cariblattoides labandeirai sp.n. from the Eocene sediments of Green River in Colorado, USA bear only two plesiomorphies, but also several significant autapomorphies within the advanced and highly derived living cockroach genus. Thus, Cariblattoides with extant occurrence in the Caribbean and South America was historically common in the Nearctic, and represents important evidence for the occurrence of derived living genera of cockroaches -50 Ma ago. Generally, the vast majority of living genera were absent during the Palaeocene, thus the diversification of most living cockroach lineages near the Palaeocene/Eocene boundary must have been extremely rapid. Females of living C. suave, the type species, have identical (sophisticated) coloration ofpronotum, but the most related living taxa are C. piraiensis and C.fontesi from Brazil (supported by phylogenetical analysis).