The genus and species Yuyuanozoon magnificissimi gen. et sp. nov., a new fossil vetulicolian, is reported from the Early Cambrian Chengjiang Fauna in Yunnan of China. It has a bipartite body plan and five pairs of cha...The genus and species Yuyuanozoon magnificissimi gen. et sp. nov., a new fossil vetulicolian, is reported from the Early Cambrian Chengjiang Fauna in Yunnan of China. It has a bipartite body plan and five pairs of chain-formed gill sacs, showing the general characteristics of Ventulicolia. However, the exclusive 'atrial cavity' and external gill observed indicate that the new form is different from those previously described as vetulicolians, probably representing a lineage developed within deuterostomes and more primitive than urochordates.展开更多
This paper gives a brief report on the new occurrence of the Early CambrianQiongzhusi' an Chengjiang fauna in the Haikou area, Kunming and its assemblage feature andthe stratigraphic section of the horizon yieldin...This paper gives a brief report on the new occurrence of the Early CambrianQiongzhusi' an Chengjiang fauna in the Haikou area, Kunming and its assemblage feature andthe stratigraphic section of the horizon yielding these fossils. Eight genera and 8 species ofTrilobitoidea are described here, embracing Leanchoilia asiatica Luo et Hu (sp. nov.), Yohoiasinensis Luo et Hu (sp. nov.), Zhongxinia speciosa Luo et Hu (gen. et sp. nov.), Xandarellaspectuculum Hou et al., Dianchia mirabilis Luo et Hu (gen. et sp. nov.), Kuamaia lata Hou,Retifacies longispinus Luo et Hu (sp. nov.) and Sinoburius lunaris Hou et al.展开更多
Well known for its abundant and extraordinary soft-tissue fossils,the Chengjiang fauna has witnessed the main phase of the Cambrian explosion and the first great congress of the ancestors of nearly all major phyla of ...Well known for its abundant and extraordinary soft-tissue fossils,the Chengjiang fauna has witnessed the main phase of the Cambrian explosion and the first great congress of the ancestors of nearly all major phyla of animals on Earth.The large-scale survey and exploration of the fauna by Chinese paleontologists,partly in collaboration with international scientists,over the past 30 years can be broadly divided into three stages.In the first decade since 1984(19841994),a large number of invertebrates,including basal animals and the protostomes of the early animal tree were discovered,but the subkingdom Deuterostomia was completely unknown.Then in the second decade(19952005),the great discoveries of various deuterostomes led to the construction of the basic framework of the Deuterostomia and then to the formation of the tripartite phylogenetic trees of early animals(TPTEA,including basal animals,protostomes and deuterostomes),for the first time.In the third decade(after 2005),the academic community began to think about the internal relationships between the formation of TPTEA and the multiepisode Cambrian explosion,leading to the new hypothesis of the three-episode Cambrian explosion.The Chengjiang fauna is important for deciphering the fauna structure,paleoecological environment,and so on.However,its core academic values mainly rest on two aspects.Firstly,The Chengjiang fauna,as the main phase witness of the Cambrian explosion,has created a nearly complete phylogenetic framework of the TPTEA on Earth for the first time.The three-phase radiation hypothesis reveals the essential connotation of the Cambrian explosion:a step-wise divergent evolution of animals,from basal to highly advanced groups,lasting about 40 million years.In the first phase,it gave birth to a bulk of basal animals(including some now extinct“animal”groups)in the latest Ediacaran,probably including some pioneer protostomes.The second phase took place in the first epoch of the Cambrian period(Terreneuvian),giving rise to the main invertebrate protostomes with a persistent prosperity of basal animals.The third phase proceeded in Cambrian Epoch 2(represented by the Chengjiang fauna),which not only maintained the prosperity of basal animals and protostomes,but also,more importantly,gave birth to all the main phyla of the subkingdom Deuterostomia.Thus,the rudimental framework of the whole TPTEA has been shaped,with the termination of the major innovation events of the Cambrian explosion.Here,we discussed the evolutionary properties of Ediacaran biota,small shelly fossils and the Chengjiang fauna in the Cambrian explosion with emphasis on the biological properties of several important animal groups.The order Myllokunmingiida is the only known oldest vertebrate,while Yunnanozoon and Haikouella are neither vertebrates nor stem-group chordates but a special group of basal deuterostomes;Cheungkongella is a credible ancestor of the urochordate and it supports the classical hypothesis on the origin of the urochordates;and the gill slits were first invented in the members of the phylum Vetulicolia to provide key information on the origin of the deuterostomes.The second core value of the Chengjiang fauna is of profound humanistic and philosophical significance:the discoveries of the‘first gill openings’,‘first brain’,‘first vertebrae’and‘first heart’provide the pivotal evidence for solving the unsolved mystery of the origin of the main basic human organs as described in Darwin’s“The Descent of Man”.In addition,the morphological and anatomical information of the Chengjiang fauna can provide important clues for a better understanding of most components of Ediacaran and Cambrian metazoans.展开更多
Twenty black shale samples, which are free from the influence of weathering, were collected from the Chengjiang Fauna-bearing horizon, central Yunnan Province, yielding a Pb-Pb isochron age of 534±60 Ma. Although...Twenty black shale samples, which are free from the influence of weathering, were collected from the Chengjiang Fauna-bearing horizon, central Yunnan Province, yielding a Pb-Pb isochron age of 534±60 Ma. Although this age is younger than both the Rb-Sr isochron age and 40Ar-39Ar age, it should represent the lower isotopic age limit of the Chengjiang Fauna.展开更多
目的系统分析滇东泥质基底澄江化石库腕足动物的生活方式。方法依据澄江化石库丰富的腕足动物资料,论证寒武纪早期泥质基底腕足动物的生活策略。结果寒武纪早期泥质基底上腕足动物的生活方式主要包括肉茎附着型(pedicle-anchoring or pe...目的系统分析滇东泥质基底澄江化石库腕足动物的生活方式。方法依据澄江化石库丰富的腕足动物资料,论证寒武纪早期泥质基底腕足动物的生活策略。结果寒武纪早期泥质基底上腕足动物的生活方式主要包括肉茎附着型(pedicle-anchoring or pedicle-attaching)、自由平躺型(free-lying)、假内栖型(qusi-infaunal)和表栖粘附型(cemented epifaunal),肉茎附着型是早寒武世腕足动物生活的最普遍的型式。结论空间生态分析显示,早寒武世滇东海盆中,腕足动物的分布已经具有一定的空间层次性,不同的腕足类型占据了不同的生态位,形成了比较复杂的空间群落结构。展开更多
本文简要报道了在昆明市海口地区下寒武统筇竹寺阶发现的澄江动物群化石,及其组合面貌和产出层位的地层剖面,重点描述了其中的三叶形虫:Leanchoilia asiatica Luo et Hu(sp. nov. ), Yohoia sinensis Luo et Hu(sp. nov. ),Zhongxinia s...本文简要报道了在昆明市海口地区下寒武统筇竹寺阶发现的澄江动物群化石,及其组合面貌和产出层位的地层剖面,重点描述了其中的三叶形虫:Leanchoilia asiatica Luo et Hu(sp. nov. ), Yohoia sinensis Luo et Hu(sp. nov. ),Zhongxinia speciosa Luo et Hu(gen. et sp. nov. ), Xandarella spectuculum Houet al. , Dianchia mirabilis Luo et Hu(gen. et sp. nov. ),Kuamaia lata Hou, Reti-facies longispinus Luo et Hu(sp. nov. ), Sinoburius lunaris Hou et al.。海口地区澄江动物群的发现,不仅扩大了澄江动物群的分布范围,也丰富了该动物群的内容。展开更多
对云南晋宁梅树村剖面下寒武统黑林铺组(原筇竹寺组)玉案山段的蠕形动物化石Sabellidites yunnanensis Luo et Zhang,S.badaowanensis Luo et Zhang的模式标本进行重新研究,通过虫体躯干表皮骨板的排列方式与产于同一剖面相同层位的古蠕...对云南晋宁梅树村剖面下寒武统黑林铺组(原筇竹寺组)玉案山段的蠕形动物化石Sabellidites yunnanensis Luo et Zhang,S.badaowanensis Luo et Zhang的模式标本进行重新研究,通过虫体躯干表皮骨板的排列方式与产于同一剖面相同层位的古蠕虫Mafangscolex sinensis Hu 2005以及Palaeoscolex sinensis Hou et Sun躯干体环上骨板排列方式进行对比,认为前3个种躯干表面骨板的排列方式完全相同,即每一体环均有5横排骨板,但与中华古蠕虫(Palaeoscolex sinensis)每一体环只有4横排大骨板的特征不相吻合。建议采用Mafangscolex yunnanensis作为前3者的属种名称,并以最早发表的Sabellidites yunnanensis Luo et Zhang作为该属的模式种。展开更多
基金supported by the National Natural Science Foundation of China(Grants 40073007,40173002,40232020)the Natural Scienee Foundation of Yunnan Province(Grant 97D007Z)
文摘The genus and species Yuyuanozoon magnificissimi gen. et sp. nov., a new fossil vetulicolian, is reported from the Early Cambrian Chengjiang Fauna in Yunnan of China. It has a bipartite body plan and five pairs of chain-formed gill sacs, showing the general characteristics of Ventulicolia. However, the exclusive 'atrial cavity' and external gill observed indicate that the new form is different from those previously described as vetulicolians, probably representing a lineage developed within deuterostomes and more primitive than urochordates.
基金This work is a contribution to Project 92D087 supported by the Yunnan Provincial Fundation for Applied Basic Researches and Project 8502004-1A supported by the fund from the Ministry of Geology and Mineral Resources for important basic researches in the
文摘This paper gives a brief report on the new occurrence of the Early CambrianQiongzhusi' an Chengjiang fauna in the Haikou area, Kunming and its assemblage feature andthe stratigraphic section of the horizon yielding these fossils. Eight genera and 8 species ofTrilobitoidea are described here, embracing Leanchoilia asiatica Luo et Hu (sp. nov.), Yohoiasinensis Luo et Hu (sp. nov.), Zhongxinia speciosa Luo et Hu (gen. et sp. nov.), Xandarellaspectuculum Hou et al., Dianchia mirabilis Luo et Hu (gen. et sp. nov.), Kuamaia lata Hou,Retifacies longispinus Luo et Hu (sp. nov.) and Sinoburius lunaris Hou et al.
基金Natural Science Foundation of China(41672009,41621003,41772010,41720104002)The Strategic Priority Research Program of Chinese Academy of Sciences(XDB18000000,XDB26000000)。
文摘Well known for its abundant and extraordinary soft-tissue fossils,the Chengjiang fauna has witnessed the main phase of the Cambrian explosion and the first great congress of the ancestors of nearly all major phyla of animals on Earth.The large-scale survey and exploration of the fauna by Chinese paleontologists,partly in collaboration with international scientists,over the past 30 years can be broadly divided into three stages.In the first decade since 1984(19841994),a large number of invertebrates,including basal animals and the protostomes of the early animal tree were discovered,but the subkingdom Deuterostomia was completely unknown.Then in the second decade(19952005),the great discoveries of various deuterostomes led to the construction of the basic framework of the Deuterostomia and then to the formation of the tripartite phylogenetic trees of early animals(TPTEA,including basal animals,protostomes and deuterostomes),for the first time.In the third decade(after 2005),the academic community began to think about the internal relationships between the formation of TPTEA and the multiepisode Cambrian explosion,leading to the new hypothesis of the three-episode Cambrian explosion.The Chengjiang fauna is important for deciphering the fauna structure,paleoecological environment,and so on.However,its core academic values mainly rest on two aspects.Firstly,The Chengjiang fauna,as the main phase witness of the Cambrian explosion,has created a nearly complete phylogenetic framework of the TPTEA on Earth for the first time.The three-phase radiation hypothesis reveals the essential connotation of the Cambrian explosion:a step-wise divergent evolution of animals,from basal to highly advanced groups,lasting about 40 million years.In the first phase,it gave birth to a bulk of basal animals(including some now extinct“animal”groups)in the latest Ediacaran,probably including some pioneer protostomes.The second phase took place in the first epoch of the Cambrian period(Terreneuvian),giving rise to the main invertebrate protostomes with a persistent prosperity of basal animals.The third phase proceeded in Cambrian Epoch 2(represented by the Chengjiang fauna),which not only maintained the prosperity of basal animals and protostomes,but also,more importantly,gave birth to all the main phyla of the subkingdom Deuterostomia.Thus,the rudimental framework of the whole TPTEA has been shaped,with the termination of the major innovation events of the Cambrian explosion.Here,we discussed the evolutionary properties of Ediacaran biota,small shelly fossils and the Chengjiang fauna in the Cambrian explosion with emphasis on the biological properties of several important animal groups.The order Myllokunmingiida is the only known oldest vertebrate,while Yunnanozoon and Haikouella are neither vertebrates nor stem-group chordates but a special group of basal deuterostomes;Cheungkongella is a credible ancestor of the urochordate and it supports the classical hypothesis on the origin of the urochordates;and the gill slits were first invented in the members of the phylum Vetulicolia to provide key information on the origin of the deuterostomes.The second core value of the Chengjiang fauna is of profound humanistic and philosophical significance:the discoveries of the‘first gill openings’,‘first brain’,‘first vertebrae’and‘first heart’provide the pivotal evidence for solving the unsolved mystery of the origin of the main basic human organs as described in Darwin’s“The Descent of Man”.In addition,the morphological and anatomical information of the Chengjiang fauna can provide important clues for a better understanding of most components of Ediacaran and Cambrian metazoans.
基金This research project was granted jointly by the National Natural Science Foundation of China(Grant No.40073001)the Key Project(97D0072)of Yunnan Provincial Foundation.
文摘Twenty black shale samples, which are free from the influence of weathering, were collected from the Chengjiang Fauna-bearing horizon, central Yunnan Province, yielding a Pb-Pb isochron age of 534±60 Ma. Although this age is younger than both the Rb-Sr isochron age and 40Ar-39Ar age, it should represent the lower isotopic age limit of the Chengjiang Fauna.
文摘澄江生物群是认识寒武纪生命大爆发的重要窗口,它的分布范围一直仅限于滇东地区,最近在贵州遵义松林牛蹄塘组底部页岩中发现有Naronia,Archotuba conoidalis,Isoxys,Perspicaris,Lingulepis,Tsunyidiscus,Scenella,水母状化石(medusiform fossil),海绵动物化石Leptomitus,类似半索动物杆壁虫Rhabdopleura化石,软舌螺类Hyolithids,宏观藻类和疑源类化石Zunyiphyton perelegans Yang et Zhao,1999,Yuknessia sp.,Longenema Ding,1996,Sphaerocongregus variabilis等。其中有一部分是属于澄江生物群分子,这是云南省外澄江生物群分子的新发现。由于生物群之下40m处还产大量海绵动物、高肌虫和藻类化石等组成的松林生物群,其上又有下寒武统明心寺组古杯动物群,因此,这一生物群的发现对研究早寒武世生物的演化也具有一定的科学意义,同时,对澄江生物群的古生态学研究也具有重要意义。
文摘目的系统分析滇东泥质基底澄江化石库腕足动物的生活方式。方法依据澄江化石库丰富的腕足动物资料,论证寒武纪早期泥质基底腕足动物的生活策略。结果寒武纪早期泥质基底上腕足动物的生活方式主要包括肉茎附着型(pedicle-anchoring or pedicle-attaching)、自由平躺型(free-lying)、假内栖型(qusi-infaunal)和表栖粘附型(cemented epifaunal),肉茎附着型是早寒武世腕足动物生活的最普遍的型式。结论空间生态分析显示,早寒武世滇东海盆中,腕足动物的分布已经具有一定的空间层次性,不同的腕足类型占据了不同的生态位,形成了比较复杂的空间群落结构。
文摘本文简要报道了在昆明市海口地区下寒武统筇竹寺阶发现的澄江动物群化石,及其组合面貌和产出层位的地层剖面,重点描述了其中的三叶形虫:Leanchoilia asiatica Luo et Hu(sp. nov. ), Yohoia sinensis Luo et Hu(sp. nov. ),Zhongxinia speciosa Luo et Hu(gen. et sp. nov. ), Xandarella spectuculum Houet al. , Dianchia mirabilis Luo et Hu(gen. et sp. nov. ),Kuamaia lata Hou, Reti-facies longispinus Luo et Hu(sp. nov. ), Sinoburius lunaris Hou et al.。海口地区澄江动物群的发现,不仅扩大了澄江动物群的分布范围,也丰富了该动物群的内容。
文摘对云南晋宁梅树村剖面下寒武统黑林铺组(原筇竹寺组)玉案山段的蠕形动物化石Sabellidites yunnanensis Luo et Zhang,S.badaowanensis Luo et Zhang的模式标本进行重新研究,通过虫体躯干表皮骨板的排列方式与产于同一剖面相同层位的古蠕虫Mafangscolex sinensis Hu 2005以及Palaeoscolex sinensis Hou et Sun躯干体环上骨板排列方式进行对比,认为前3个种躯干表面骨板的排列方式完全相同,即每一体环均有5横排骨板,但与中华古蠕虫(Palaeoscolex sinensis)每一体环只有4横排大骨板的特征不相吻合。建议采用Mafangscolex yunnanensis作为前3者的属种名称,并以最早发表的Sabellidites yunnanensis Luo et Zhang作为该属的模式种。
