Clematis sect. Atragene is revised in this paper based on the examination of a large number of herbarium specimens, extensive field observations, and morphometric analyses. Brief taxonomic history and geographical dis...Clematis sect. Atragene is revised in this paper based on the examination of a large number of herbarium specimens, extensive field observations, and morphometric analyses. Brief taxonomic history and geographical distribution of the section are given, the relationships among the species are discussed, and the evolutionary trends of some characters in the section are evaluated. The staminodes of the plants in this section may have evolved from the outer stamens with petaloid filaments and gradually disappearing anthers. Subsequently, they may have evolved in two different ways. One possibility is that the staminodes elongate and become lanceolate, as long as sepals, and their apices turn into attenuate. The other is that the staminodes are spathulate, but not elongating, as long as stamens, and their apices turn into retuse from obtuse and rounded. The evolutionary trend of sepals may be from thin to thick in texture, and the veins from non-prominent to prominent. As a result, five new series are established and nine species, two subspecies and nine varieties （including three new ranks） are recognized in this section. An identification key is provided, and each taxon is described and illustrated. Clematis sibirica and Clematis ochotensis are treated as subspecies of Clematis alpina due to their subtle differences and lack of, or few, overlapping distributions. Clematis fusijamana and Clematisfauriei are recognized as varieties of C. alpina ssp. ochotensis for the continuous variation of the velutinous strips on the sepal margins. Clematis iliensis is treated as variety of C. alpina ssp. sibirica for the continuous variation of leaf division types. Extensive variations in sepal color and basal caruncle size support degrading Clematis chiisanensis as a variety of Clematis koreana. The North American ser. Occientales may be primitive, whereas ser. Macropetalae may be the most advanced taxon in this section. Ser. Alpinae and ser. Koreanae are closely related to each other. However, the systematic position of ser. Tomentosae cannot be determined based on morphological characters alone in the present study.展开更多
In this paper Clematis sect. Aspidanthera s.l. is revised. Seventy_ two species and 15 varieties are recognized. They are keyed, described, and in most cases illustrated, and are classified into six subsections. Of th...In this paper Clematis sect. Aspidanthera s.l. is revised. Seventy_ two species and 15 varieties are recognized. They are keyed, described, and in most cases illustrated, and are classified into six subsections. Of them, on e series, five species, and two varieties are described as new, and two new rank s are proposed. Brief taxonomic history and geographical distribution of the sec tion are given, the evolutionary trends of several important morphological characters observed are e numerated, and the relationships of its subsections are discussed. As a resul t, subsect. Dioicae, which shows striking resemblance in both habit an d floral structure to sect. Clematis subsect. Clematis, is regar ded as t he primitive group, while subsect. Insidiosae, bearing pistillate flower wit h erect sepals and no staminodes, is regarded as the advanced group within the s ection.展开更多
基金supported by the National Natural Science Foundation of China(Grant No.30870146).
文摘Clematis sect. Atragene is revised in this paper based on the examination of a large number of herbarium specimens, extensive field observations, and morphometric analyses. Brief taxonomic history and geographical distribution of the section are given, the relationships among the species are discussed, and the evolutionary trends of some characters in the section are evaluated. The staminodes of the plants in this section may have evolved from the outer stamens with petaloid filaments and gradually disappearing anthers. Subsequently, they may have evolved in two different ways. One possibility is that the staminodes elongate and become lanceolate, as long as sepals, and their apices turn into attenuate. The other is that the staminodes are spathulate, but not elongating, as long as stamens, and their apices turn into retuse from obtuse and rounded. The evolutionary trend of sepals may be from thin to thick in texture, and the veins from non-prominent to prominent. As a result, five new series are established and nine species, two subspecies and nine varieties （including three new ranks） are recognized in this section. An identification key is provided, and each taxon is described and illustrated. Clematis sibirica and Clematis ochotensis are treated as subspecies of Clematis alpina due to their subtle differences and lack of, or few, overlapping distributions. Clematis fusijamana and Clematisfauriei are recognized as varieties of C. alpina ssp. ochotensis for the continuous variation of the velutinous strips on the sepal margins. Clematis iliensis is treated as variety of C. alpina ssp. sibirica for the continuous variation of leaf division types. Extensive variations in sepal color and basal caruncle size support degrading Clematis chiisanensis as a variety of Clematis koreana. The North American ser. Occientales may be primitive, whereas ser. Macropetalae may be the most advanced taxon in this section. Ser. Alpinae and ser. Koreanae are closely related to each other. However, the systematic position of ser. Tomentosae cannot be determined based on morphological characters alone in the present study.
文摘In this paper Clematis sect. Aspidanthera s.l. is revised. Seventy_ two species and 15 varieties are recognized. They are keyed, described, and in most cases illustrated, and are classified into six subsections. Of them, on e series, five species, and two varieties are described as new, and two new rank s are proposed. Brief taxonomic history and geographical distribution of the sec tion are given, the evolutionary trends of several important morphological characters observed are e numerated, and the relationships of its subsections are discussed. As a resul t, subsect. Dioicae, which shows striking resemblance in both habit an d floral structure to sect. Clematis subsect. Clematis, is regar ded as t he primitive group, while subsect. Insidiosae, bearing pistillate flower wit h erect sepals and no staminodes, is regarded as the advanced group within the s ection.
基金Supported by the National Natural Science Foundationof China(GrantNos.30070059,39570051)and the Special Projectfor Taxonomicand Floristic Researches from the Chinese Academy of Sciences.
基金Supported by the National Natural Science Foundation of China, Grant No. 30470126, and the Special Project for Taxonomic and Floristic Researches from the Chinese Academy of Sciences.
