基金Funding: This study was funded by the National Natural Science Foundation of China (81260488 and 81560669), KeyLab Construction Project of the Educational Department of Guizhou Province (Project No. Guizhou EducationCooperation KY[2014]212) , Science Inovative Talent Team for Medicinal Insect Research and Development in Zunyi(Zunyi shi ke he 2015-40) and Modernization of Traditional Chinese Medicine in Guizhou Province high-tech researchand development projects (Qian ke he ZY 2012-2009).