New Jurassic Fossil True Bugs of the Pachymeridiidae(Hemiptera:Pentatomomorpha)from Northeast China

Four new fossil genera and species of true bugs from the family Pachymeridiidae, Beipiaocoris multifurcus, Bellicoris mirabilis, Nitoculus regiUus, and Viriosinervis stolidus, are described. New specimens were collected from the Middle and Upper Jurassic non-marine sedimentary strata from the Jiulongshan and Yixian Formations of northeast China. The species Karatavocoris asiatica Becker-Migdisova, 1963, which was considered to be a member of the family Coreidae is transferred to the Pachymeridiidae. A new map of all known and newly discovered fossil pachymeridiid localities is given. The diagnosis of the family is modified.

Late Mesozoic Chresmodids with Forewing from Inner Mongolia, China (Polyneoptera: Archaeorthoptera)

Three new species of Chresmodidae are described. They belong to two different genera： Chresmoda multinervis sp. nov., Chresmoda shihi sp. nov. and Jurachresmoda sanyica sp. nov.. A study of the forewing venation is provided. Longitudinal veins are mostly parallel; costal area is broad at the basal and narrowing distally; ScP is probably extending close to the wing apex; R is parallel at length to ScP; MA is running close to R, with two long parallel branches; MA branched-off basally; while MP is not branched until the middle of the wing. However, the important basal part of MP and CuP is poorly preserved. In addition, unique fringing hairs on the mid legs of C. shihi sp. nov. and J. sanyica sp. nov. are described and compared with Jurachresmoda gaskelli Zhang, Ren and Shih, 2008.

Two New Fossil Lacewing Species from the Middle Jurassic of Inner Mongolia,China(Neuroptera:Grammolingiidae)

Two new species of lacewings are described and illustrated from the Jiulongshan Formation （Middle Jurassic） of Inner Mongolia, China： Leptolingia calonervis sp. nov. and Litholingia ptesa sp.nov. （Grammolingiidae）. The species can be distinguished by the following features： the outermost branch of MP2 on the hind wing did not reach the margin of wing in Leptolingia calonervis sp. nov.;the second branch of Rs forked deeply while other branches of Rs forked much later in Litholingia ptesa sp. nov..

New Species of Cimbrophlebiidae(Insecta: Mecoptera) from the Middle Jurassic of Northeastern China

Four new species of three genera in the family Cimbrophlebiidae, Telobittacus decorus sp.nov., Bellicimbrophlebia heteroneura sp.nov., Cimbrophlebia amoena sp.nov., and Cimbrophlebia gracilenta sp.nov., are described and illustrated based on eleven well-preserved fossil specimens.These specimens were collected from the latest Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China.This is the first record of the genus Cimbrophlebia in the Jurassic extending its age from the Lower Cretaceous to the Middle Jurassic.Due to same venational characters, we propose that an undescribed fossil specimen of Cimbrophlebiidae from the Jurassic of Germany is attributed to C.gracilenta sp.nov.Our new findings demonstrate an evolutionary trend of venational change from 185 to 50 Ma that the level of the bifurcation of Rs is gradually moving toward the base of wing in reference to the bifurcation of M.

Two New Wasps (Hymenoptera:Stephanoidea:Ephialtitidae) from the Middle Jurassic of China

Two new species of wasps belonging to the family Ephialtitidae,Praeproapocritusflexus sp.nov.and Proapocritus parallelus sp.nov.,from the Middle Jurassic Jiulongshan Formation of southeastern Inner Mongolia,China,are described and illustrated.Praeproapocritus vulgates Rasnitsyn ＆ Zhang,2010 and six species of Proapocritus,P.densipediculus,P.sculptus,P.longantennatus,P.formosus,P.atropus,and P.elegans,all described by Rasnitsyn and Zhang in 2010,are first record from the Middle Jurassic of China.These two new findings supplement the morphological understanding of ephialtitids in the Middle Jurassic of China and broaden the diversity of ephialtitids in their by-gone ecosystems.

First Record of Fossil Mesocupes from China(Coleoptera:Archostemata:Cupedidae)

Three new fossil species of the genus Mesocupes of fossil cupedids, M. angustilabialis sp. nov., M. latilabialis sp. nov. and M. collaris sp. nov., are described from the Jiulongshan Formation of Inner Mongolia, China. These new species are the first three records of fossil Mesocupes in China as well as the oldest ones in the world now. This finding also extends the geographical distribution of this genus from Central to East Asia. In addition, based on the ratio of the length of the last to the penultimate abdominal ventrites of new beetles distinctly lower than that of the species from the Karabastau Formation, the age of Daohugou fossil-bearing beds might be older than that of Karatau assemblage and consequently of Middle Jurassic.

New Genus and Species of Rhyacophilidae (Insecta:Trichoptera) from the Middle Jurassic of China

One new genus with a new fossil species,Declinimodus setulosus gen.et sp.nov.which was found in the Middle Jurassic Jiulongshan Formation of Daohugou,Inner Mongolia,China,are described and illustrated.The new species is assigned to the Rhyacophilidae according to buttonshaped of the 2nd segment of maxillary palp,and the forked R1 （in the forewing,located near apex）.These new discovery are the earliest fossil records of Rhyacophilidae,which providing important evident for the study of the origin and early evolution of Trichoptera.

A New Fossil Scorpionfly(Insecta:Mecoptera:Holcorpidae) with Extremely Elongate Male Genitalia from Northeastern China

Since Holcorpa maculosa was first described in 1878, it has attracted lots of attention due to its extremely elongate abdominal segments from the sixth to the eighth. Fossil records of family Holcorpidae are very rare, comprising two species in one genus represented by three fossil specimens, resulting in limited knowledge on the structure and function of holcorpid＇s unique male organ, as well as its evolutionary relationship with other scorpionflies. Herein we describe a new genus with a new species, Conicholcorpa stigmosa gen. et sp. nov., of the Holcorpidae from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Inner Mongolia, China. This is the first holcorpid documented from the Daohugou locality, albeit two previously reported species with extremely elongate male organs but deemed as Family Incertae sedis. C. stigmosa gen. et sp. nov. is the earliest holcorpid hitherto, extending its existence from the Early Eocene to the Middle Jurassic by 114 million years and providing one more early case of presenting exaggerated male body parts for sexual display and/or selection. Our analysis supports the sister relationship between Holcorpidae and Orthophlebiidae. Our results not only enhance our knowledge of the morphology of Holcorpidae but also elucidate the relationship of known species of this family.

New Species of Cretolimonia and Mesotipula(Diptera:Limoniidae) from the Middle Jurassic of Northeastern China

Two new species, Cretolimonia excelsa sp. nov. and Mesotipula gloriosa sp. nov., in Limoniidae of Diptera are described and illustrated, based on three fossil specimens with bodies and complete wings. These specimens were collected from the latest Middle Jurassic Jiulongshan Formation at Daohugou, Inner Mongolia, China. These are the first fossil records of these two genera from China.

New fossil evidence of the early diversification of scarabs： AIIoioscarabaeus cheni （Coleoptera： Scarabaeoidea） from the Middle Jurassic of Inner Mongolia, China

Scarabaeoidea are known from the Lower Jurassic and may have originated in the Triassic based on fossil evidence and phylogenetic research. However, the early diversification of Scarabaeoidea remains unclear due to the lack of high-quality fossil evidence. Here we describe an exceptionally well-preserved new fossil of Scarabaeoidea, Alloioscarabaeus cheni gen. et sp. nov from the Middle Jurassic Jiulongshan Formation of Inner Mongolia, China. Based on a morphometric analysis using 17 landmarks of the hind wing of Alloioscarabaeus and 10 scarabaeoid families, we found that Alloioscarabaeus cheni gen. et sp. nov clearly does not belong to any of the known scarabaeoid families and, consequently, is a new family, Alloioscarabaeidae fam. nov., was erected. The discovery of Alloioscarabaeus brought further evidence for the early diversification of major scarab lineages which could allow more detail in the palaeobiogeography of the Scarabaeoidea and Northeast of China which might be one of the originating places or an important radiation place during the evolution of Scarabaeoidea. Alloioscarabaeidae were very likely not good diggers and might have fed on decaying organic materials. Based on the evidence we have now, we tend to believe that most families and some subfamilies of Scarabaeoidea were present in the Jurassic period.