Impacts of livelihood assets on livelihood security in drought-prone Gamo lowlands of southwest Ethiopia

Livelihood assets are a matter of high concern for secured survival.Drought-prone Gamo lowland households have differential access to livelihood resources which indicates the varying capacity of resisting to shocks.The main objective of this study is to explore the impacts of livelihood assets on livelihood security in the drought-prone Gamo lowlands.Multistage sampling procedures were employed to select the study sites and sample respondents.Primary data of households’capital assets and livelihood security status were produced from 285 survey households,agricultural experts,key informants,focus group discussants,and field observation through transect walks.Descriptive and inferential statistics were used to analyze quantitative data,whereas discussions and annotations were employed for analyzing qualitative data.The Sustainable Livelihoods Framework is used with modifications to schematize the study conceptually.The findings indicated that the study households possessed combinations of livelihood resources differentially.Financial and natural capitals were found to be the most deficient and better-accessed capitals,respectively.The study also showed that lowland residents’access to assets has significant indications of livelihood security.Households’poor access to assets such as financial,information,and social capital demands raised attention of the concerned stakeholders and policy debates in the drought-prone rural setup.Hence,it has been concluded that the more assets are accessed,the stronger the capacity of the households to resist shocks,and better the livelihood security.Accordingly,enhancing people’s access to multiple livelihood assets is suggested to sustainably secure livelihoods.

FLOOD SIMULATION IN THE COASTAL LOWLANDS OF NORTHERN JIANGSU,CHINA

The coastal lowlands in northern Jiangsu Province, China, is characterized by low relief and low water surface gradient in streams. The flooding stages are commonly lower than the high-tide level and the tidal range is relatively small. All these factors prohibit rapid discharge of floodwater and make the flood regimes here extremely sensitive to the sea level rise. The 1991 Plum Rain was examined, when precipitation was significantly higher than normal. The one-dimensional open-channel non-conservative flow model was used to simulate the stream flow in the channel network of studied area. GIS technology was used to delineate the maximum in undated area by analyzing the DEMs.

The Linkage between Natural Vegetation, Water Dynamics and Pyrite （FeS2） Oxidation in Tidal Lowlands

The research aimed to analyze the linkage between natural vegetation, water dynamics and pyrite （FeS2） oxidation in tidal lowlands. The research was carried out in tidal lowland Pulau Rimau, South Sumatra from February to December 2010. The field observations are done by exploring several transect on land units. The field description refers to Soil Survey Staff. Water and soil samples were taken from selected key areas for laboratory analyses. The vegetation data were collected by making sample plots placed on each vegetation type with plot sizes 10 m × 10 m for secondary forests and 5 m × 5 m for shrubs and grass. The observations of surface water level were done during the river receding with units of meter above sea level （m.asl）. The results shows that pyrite formation is largely determined by the availability of natural vegetation as S （sulfur） donors, climate and uncontrolled water balance and supporting faunas such as crabs and mud shrimp. Climate and water balance as well as supporting faunas is the main supporting factors to accelerate the process of formation pyrite. Oxidized pyrite increases soil pH thus toxic to fish, arable soils, plant growth, disturbing the water quality and soil nutrient availability. Oxidized pyrite is predominantly accelerated by the dynamics of river water and disturbed natural vegetation by human activities, and the pyrite oxidation management approach is divided into three main components of technologies, namely water management, land management and commodity management.

Traditional Lowlands Water Management in Dano, South-Western Burkina Faso

Lowland water resources management represents a challenge of the future that calls any community. Irrigated crops are grown in some areas of Burkina Faso, others are limited by a lack of irrigation infrastructure. Due to limited crop irrigation, crops and the associated populations dependent on them, depend on rain and on climatic factors. Thus, there is a need to understand and implement traditional mechanisms for managing lowland water in Dano, where climatic and geological conditions provide a sustained source of water. Here, I use a literature review combined with field work and interviews/questionnaires to estimate the potential exploitable plains to 16,056 ha or 24% of the communal area. Management mechanisms and traditional operating systems of lowland waters were clear, which helped to set the technological level of farmers, in partial control of water management.

Assessing Silicon Availability in Soils of Rice-Growing Lowlands and Neighboring Uplands in Benin and Nigeria

Silicon(Si) is known as a beneficial nutrient in the cultivation of rice, playing a key role in photosynthesis enhancement, lodging resistance and tolerance to various environmental stress. The present study aimed to examine available Si content in both lowland soils(n = 29) and neighboring upland soils(n = 21) collected from Benin and Nigeria and to evaluate the validity of the assessment results through a pot experiment. Our results revealed that the acetate-buffer method predicted Si concentration in rice straw at the harvest stage(R^2 = 0.68, P < 0.01) better than the anaerobic-incubation method(R2 = 0.31, P > 0.05), and 76% of the uplands and 38% of the lowlands were deficient(< 50 mg/kg) in acetate-buffer soluble Si. These findings suggest that the Si-deficiency soils prevail across the study area, making rice plants starved for Si and prone to environmental stress.

Impact of Smart Valley on Soil Moisture Content and Rice Yield in Some Lowlands in Burkina Faso

To reduce the impact of rainfall variability on lowland rice yields, Burkina Faso state develops lowlands for small rice farmers. However, the high cost of these infrastructures makes impossible to duplicate them to satisfy the needs which are enormous. The Smart-Valley technology which is actually popularized in certain coastal countries of West Africa would therefore be a boon to increase the productivity of the Sudanese lowlands if it well regulates runoff. The object of this study was therefore to know if smart valley technology could increase soil moisture in order to mitigate the impact of drought’s pockets on rice cultivation in the Sudanese lowlands. The experiment takes place in three lowlands during the rainy seasons 2018 and 2019. The climatic data comes from the meteorological stations in the study areas as well as those installed on the sites. The infiltration measurements were carried out using the double Muntz ring. The soil moisture measurement device consisted of a smart valley area of 5 ha and an undeveloped area of 5 ha per site. Sixteen tubes were installed per lowland allowing the humidity to be measured at a depth of 10, 20, 30, 40 cm using a probe. Four rice varieties, Orylux6, FKR62N, FKR19 and FKR64 were tested on plots of 0.25 ha per variety in the smart valley and undeveloped parts. The results showed that the humidity level was 12% higher in the smart-valley plots throughout the cycle compared to the unmanaged area. In addition, humidity decreases rapidly in unmanaged plots as rain becomes increasingly scarce. Finally, the smart-valley development allowed an average increase in rice yields of 21% compared to the average yield of undeveloped plots.

Woody plant taxonomic,functional,and phylogenetic diversity decrease along elevational gradients in Andean tropical montane forests:Environmental filtering and arrival of temperate taxa

Mountains are paramount for exploring biodiversity patterns due to the mosaic of topographies and climates encompassed over short distances.Biodiversity research has traditionally focused on taxonomic diversity when investigating changes along elevational gradients,but other facets should be considered.For first time,we simultaneously assessed elevational trends in taxonomic,functional,and phylogenetic diversity of woody plants in Andean tropical montane forests and explored their underlying ecological and evolutionary causes.This investigation covered four transects(traversing ca.2200 m a.s.l.) encompassing 114 plots of 0.1 ha across a broad latitudinal range(ca.10°).Using Hill numbers to quantify abundance-based diversity among 37,869 individuals we observed a consistent decrease in taxonomic,functional,and phylogenetic diversity as elevation increased,although the decrease was less pronounced for higher Hill orders.The exception was a slight increase in phylogenetic diversity when dominant species were over-weighted.The decrease in taxonomic and functional diversity might be attributed to an environmental filtering process towards highlands,where the increasingly harsher conditions exclude species and functional strategies.Besides,the differences in steepness decrease between Hill orders suggest that rare species disproportionately contribute to functional diversity.For phylogenetic diversity the shifting elevational trend between Hill orders indicates a greater than previously considered influence in central Andean highlands of tropical lowlands originated species with strong niche conservatism relative to distantly related temperate lineages.This could be explained by a decreasing presence and abundance of temperate,extratropical taxa towards the central Andes relative to northern or southern Andes,where they are more prevalent.

Temporal-spatial changes in the belowground bud bank in interdune lowlands of an active sand dune ecosystem in northeastern China

Aims The belowground bud bank plays an important role in vegetation restoration of sand dune ecosystems in semi-arid regions.However,few studies have focused on the temporal-spatial changes of belowground bud banks in interdune lowlands.Methods The size and composition of belowground bud bank in five interdune lowlands with different sizes were investigated for one growing season to determine the temporal and spatial changes in belowground bud bank.Important Findings Total bud bank density was the highest in the medium-sized interdune lowland as was tiller bud density.The density of stem-base buds exhibited an opposite trend while rhizome bud density did not change with interdune lowland size.There was a significant seasonal change in the bud bank size.The total bud density peaked in August and was the lowest in October.A similar trend was found for rhizome bud density,whereas the density of stem-base buds showed an opposite trend,and tiller bud density did not change signifiesntly during the growing season.We conclude that the belowground bud bank density is changed with interdune lowland size and season.These results contribute to the understanding of adaptive strategies of plants growing in active dune ecosystems and provide pointers for adopting effective measures to restore and conserve dune vegetation in semi-arid regions.

Heat-tolerant maize for rainfed hot,dry environments in the lowland tropics:From breeding to improved seed delivery

Climate change-induced heat stress combines two challenges:high day-and nighttime temperatures,and physiological water deficit due to demand-side drought caused by increase in vapor-pressure deficit.It is one of the major factors in low productivity of maize in rainfed stress-prone environments in South Asia,affecting a large population of smallholder farmers who depend on maize for their sustenance and livelihoods.The International Maize and Wheat Improvement Center(CIMMYT)maize program in Asia,in partnership with public-sector maize research institutes and private-sector seed companies in South Asian countries,is implementing an intensive initiative for developing and deploying heat-tolerant maize that combines high yield potential with resilience to heat and drought stresses.With the integration of novel breeding tools and methods,including genomics-assisted breeding,doubled haploidy,fieldbased precision phenotyping,and trait-based selection,new maize germplasm with increased tolerance to heat stress is being developed for the South Asian tropics.Over a decade of concerted effort has resulted in the successful development and release of 20 high-yielding heat-tolerant maize hybrids in CIMMYT genetic backgrounds.Via public–private partnerships,eight hybrids are presently being deployed on over 50,000 ha in South Asian countries,including Bangladesh,Bhutan,India,Nepal,and Pakistan.

Projecting future precipitation change across the semi-arid Borana lowland,southern Ethiopia

Climate change caused by past,current,and future greenhouse gas emissions has become a major concern for scientists in the field in many countries and regions of the world.This study modelled future precipitation change by downscaling a set of large-scale climate predictor variables(predictors)from the second generation Canadian Earth System Model(CanESM2)under two Representative Concentration Pathway(RCP)emission scenarios(RCP4.5 and RCP8.5)in the semi-arid Borana lowland,southern Ethiopia.The Statistical DownScaling Model(SDSM)4.2.9 was employed to downscale and project future precipitation change in the middle(2036-2065;2050s)and far(2066-2095;2080s)future at the local scale.Historical precipitation observations from eight meteorological stations stretching from 1981 to 1995 and 1996 to 2005 were used for the model calibration and validation,respectively,and the time period of 1981-2018 was considered and used as the baseline period to analyze future precipitation change.The results revealed that the surface-specific humidity and the geopotential height at 500 hPa were the preferred large-scale predictors.Compared to the middle future(2050s),precipitation showed a much greater increase in the far future(2080s)under both RCP4.5 and RCP8.5 scenarios at all meteorological stations(except Teletele and Dillo stations).At Teltele station,the projected annual precipitation will decrease by 26.53%(2050s)and 39.45%(2080s)under RCP4.5 scenario,and 34.99%(2050s)and 60.62%(2080s)under RCP8.5 scenario.Seasonally,the main rainy period would shift from spring(March to May)to autumn(September to November)at Dehas,Dire,Moyale,and Teltele stations,but for Arero and Yabelo stations,spring would consistently receive more precipitation than autumn.It can be concluded that future precipitation in the semi-arid Borana lowland is predicted to differ under the two climate scenarios(RCP4.5 and RCP8.5),showing an increasing trend at most meteorological stations.This information could be helpful for policymakers to design adaptation plans in water resources management,and we suggest that the government should give more attention to improve early warning systems in drought-prone areas by providing dependable climate forecast information as early as possible.

Assessing Suitability of Irrigation Scheduling Decision Support Systems for Lowland Rice Farmers in Sub-Saharan Africa—A Review

Irrigation in lowland rice production systems in Sub-Saharan Africa (SSA) is mainly based on traditional surface irrigation methods with continuous flooding practices. This irrigation method ends up using a lot more water that would have otherwise been used to open more land and be used in other water-requiring sectors. Various studies suggest Alternate Wetting and Drying (AWD) as an alternative practice for water management that reduces water use without significantly affecting yield. However, this practice has not been well adopted by the farmers despite its significant benefits of reduced total water use. Improving the adoption of AWD using irrigation Decision Support Systems (DSSs) helps the farmer on two fronts;to know “how much water to apply” and “when to irrigate”, which is very critical in maximizing productivity. This paper reviews the applicability of DSSs using AWD in lowland rice production systems in Sub-Saharan Africa.

Species composition and diversity,and carbon stock in a dune ecosystem in the Horqin Sandy Land of northern China

In this study, we determined carbon allocation and carbon stocks in the plant-soil system of different dune ecosystems in northeastern China. We quantified the species composition, above and below-ground bio masses, and carbon stocks of three dune types （i.e. active dunes, semi-stabilized dunes and stabilized dunes） and their corresponding inter-dune lowlands （i.e. interdune lowlands of active dunes, interdune lowlands of semi-stabilized dunes and interdune lowlands of stabilized dunes） in the Horqin Sandy Land. The results showed that the succession series on interdune lowlands of the Horqin Sandy Land confirmed differences in species composition of the various dune types. Aboveground carbon （AGC） on the interdune lowlands of semi-stabilized dunes （33.04 g C/m2） was greater （P〈0.05） than that on the interdune lowlands of active dunes （10.73 g C/m2）. At the same time, the different dune types did not show any significant differences （/：〉〉0.05） in belowground plant carbon （BGC）. However, the percentage of plant BGC in interdune lowlands of active dunes （81.5%） was significantly higher （P〈0.05） than that in the interdune lowlands of semi-stabilized dunes （58.9%）. The predominant carbon pool in the study dune ecosystem was in the soil. It accounted for 95% to 99% of total carbon storage. Soil organic carbon （SOC） was at least 55% greater （P〈0.05） in the interdunes than in the dunes. Stabilized dunes showed at least a 37% greater （P〈0.05） SOC content than active dunes up to a 1-m soil depth. Meanwhile, SOC content of interdune lowlands of semi-stabilized dunes was greater （P〈0.05） than that of interdune lowlands of active dunes only up to a 20-cm soil depth. The dune ecosystem showed a great potential to store carbon when interdune lowlands of active dunes were conversed to interdune lowlands of semi-stabilized dunes, which stored up to twice as much carbon per unit volume as interdune lowlands of active dunes.

Engineering Strategies on Flood Control in Middle Reach of Yangtze River, China

Flood disaster has been a serious hidden danger since the ancient time. The essential cause for the fact that floods have not been eliminated for hundreds of years is that time honored strategies do not suit the cases of flood prevention. In the view of geological environmental analyses of flood formation and from the synthesis of experiences gained in flood control in the past hundreds of years, sluggish draining of flood, silt sedimentation in channel and building levee blindly constitute the main cause of intractable flood for a long time in the middle reach of the Yangtze River. Draining away silt and water is the only way to stamping out flood disaster. Opening up artificial waterways for flood diversion, draining away the silt of channel into the polders, and storing the flood water are important engineering measures for the flood control and damage reduction.

Land-use Changes Alter Energy and Water Balances on an African Brachiaria Pasture Replacing a Native Savanna in the Orinoco llanos

The seasonal changes in the energy balance after the substitution of a herbaceous savanna by a Brachiaria field located in the Orinoco lowlands were assessed over an entire year using the eddy covariance technique. Simultaneously, an herbaceous savanna was monitored as a control. This work provides evidence that the vegetation replacement lead to different patterns of energy and water balance. The seasonal trends of the latent heat flux (λE) to available energy (Ra) ratio tended to decrease as senescence increased due to seasonal influence of air humidity mole fraction deficit and soil water content on leaf area index (LAI) and surface conductance (gs). Therefore, the partitioning of the available energy depended on both climatological (i.e., solar radiation, volumetric soil water content and air humidity mole fraction deficit) and biological variables (i.e., conductance behavior and LAI) which were stress-induced. For the wet season, the seasonally averaged daily λE in the Brachiaria field (i.e., 0.8 ± 0.1 mm d^-1) was 1.3-fold higher than that in the herbaceous savanna (i.e., 0.6 ± 0.1 mm d-1)(Mann-Whitney U-test). For the dry season, the value was 2.7 ± 0.6 and 2.2 ± 0.4 mm d-1, respectively, these means values were not significantly different. In the Brachiaria and herbaceous savanna stands, the annual evapotranspiration was 731 and 594 mm year^-1, respectively, and the annual ratio of evapotranspiration to precipitation was 0.52 to 0.42 respectively. In Brachiaria field, the deep drainage was relatively lower (43% of total precipitation) than that in the herbaceous savanna stand (53%) leaving a similar amount of water to increase soil storage. The current shift in land cover decrease deep drainage and increased λE by water uptake from a pasture with high belowground phytomass and LAI.

基于DSP和FPGA的智能信号源设计

针对传统信号发生器源输出信号的频率和幅值不能快速变化的不足,提出了一种基于TMS320F28335和Cyclone Ⅱ系列FPGA以及高精度D/A的设计方案。设计中通过消除累积误差来提高频率精度,并利用Lowland电流源实现了小信号的远距离传输。实验证明:该信号频率和幅值精度满足指标要求,解决了传统信号发生器不能输出波形组合的不足。现已成功运用于某型号激光接收机的测试。

Soil Resources and Land Use in Tropical Asia

Tropical Asia is a region comprising South and Southeast Asia and under strong influence of the Asianmonsoon climate. It is characterized by an extremely high population density and by high land use intensity.Paddy rice cultivation is the most important form of agriculture in the greater part of the region. Soilresources of tropical Asia have a specific feature in comparison with tropical Africa and America. Ultisolsdominate in uplands, and lowland soils like Inceptisols and Histosols are relatively abundant. The latterpoint is made clearer if we take the landforms of the region with a vast extent of lowlands into consideration.Geologically, tropical Asia with the Himalayan orogeny and active volcanism exhibits a conspicuous contrastto tropical Africa and America with the dominance of the shield structure. This along with the monsoonclimate should have determined the basic features of landforms and soil, and accordingly all the agriculturaland social characteristics of tropical Asia today. Although paddy rice cultivation in the lowland is highlysustainable, upland cultivation in extensive Ultisol areas tends to be handicapped by low fertility and higherodibility of the soil, resulting in low sustainability. Land shortage is compelling people to exploit slopelandsin hills and mountains, on the one hand, and thus far unutilized coastal lowlands, on the other. Both ofthese new reclamations are facing to serious land degradation problems today. Tropical Asia will continueto be the most densely populated region of the world with ever-increasing population. In order to meet theincreasing food demand lowland rice cultivation should be intensified by the infrastructure development toease the stresses on slopelands and vulnerable coastal lowlands. At the same time, upland crop productionin Ultisol areas should be stabilized and enhanced, providing integrated nutrient management and measuresfor soil conservation.

Germplasm Innovation of Heat Tolerance in Rice for Irrigated Lowland Conditions in the Philippines

Heat-tolerant varieties, such as N22 and Dular, which were used in this study, usually have low yield potential and undesirable plant characteristics but combining them with high yielding and improved rice varieties, new heat-tolerant rice genotypes with high yield potential can be achieved. In this study, phenotyping and selecting desirable materials from various crosses were performed under high temperature conditions during the reproductive stage. Screening was performed in the field and glasshouse to select individuals with heat tolerance and high yield potential. Several advanced breeding lines from Gayabyeo/N22 cross produced desirable individuals with heat tolerance, resistance to pests and diseases, and high yield potential. The genetic variation in percent sterility among the selected backcross populations grown in high temperature environments showed that large number of plants can be identified and selected with lower percent sterility.

Rainforests north of the Tropic of Cancer: Physiognomy, floristics and diversity in ‘lowland rainforests’ of Meghalaya, India

The lowland rainforests of Meghalaya, India represent the westernmost limit of the rainforests north of the Tropic of Cancer. These forests, on the Shillong plateau, are akin to Whitmore's ‘tropical lowland evergreen rainforest' formation and exhibit striking similarities and conspicuous differences with the equatorial rainforests in Asia-Pacific as well as tropical seasonal rainforests in southwestern China near the Tropic of Cancer. We found these common attributes of the rainforests in Meghalaya: familial composition with predominance of Euphorbiaceae, Lauraceae, Meliaceae, Moraceae, Myrsiticaceae,Myrtaceae and Rubiaceae; deciduousness in evergreen physiognomy; dominance of mega-and mesophanerophytic life-forms; abundance of species with low frequency of occurrence(rare and aggregated species); low proportional abundance of the abundant species; and truncated lognormal abundance distribution. The levels of stand density and stand basal area were comparable with seasonal rainforests in southwestern China, but were lower than equatorial rainforests. Tropical Asian species predominated flora, commanding 95% of the abundance. The differences include overall low stature(height) of the forest, inconspicuous stratification in canopy, fewer species and individuals of liana, thicker understory,higher proportion of rare species, absence of locally endemic species and relatively greater dominance of Fagaceae and Theaceae. The richness of species per hectare(S) was considerably lower at higher latitudes in Meghalaya than in equatorial rainforests, but was comparable with seasonal rainforests. Shannon's diversity index(H’=4.40 nats for ≥10 cm gbh and 4.25 nats for ≥30 cm gbh) was lower on higher latitudes in Meghalaya in comparison to species-rich equatorial rainforests, but it was the highest among all lowland rainforests near the Tropic of Cancer.

Longitudinal observation of intraocular pressure variations with acute altitude changes

BACKGROUND Higher intraocular pressure(IOP)is a major risk factor for developing glaucoma,and the leading cause of irreversible blindness worldwide.High altitude(HA)may be involved in IOP,but the reported results were conflicting.Ascent to HA directly by plane from low altitude regions is an acute,effortless exposure.However,the effects of such exposure to different altitudes on IOP have rarely been reported.AIM To investigate changes in IOP after rapid effortless exposure to HA in stages and compare it with systemic parameters.METHODS Fifty-eight healthy subjects(116 eyes)were divided into three groups:17 lowaltitude(LA)residents[44 m above sea level(ASL)],22 HA residents(2261 m ASL)and 19 very HA(VHA)residents(3750 m ASL).The LA group flew to HA first.Three days later,they flew with the HA group to VHA where both groups stayed for 2 d.Then,the LA group flew back to HA and stayed for 1 d before flying back to 44 m.IOP,oxygen saturation(SpO2)and pulse rate were measured.The linear mixed model was used to compare repeated measurements.RESULTS IOP in the LA group significantly decreased from 18.41±2.40 mmHg at 44 m to 13.60±3.68 mmHg at 2261 m ASL(P<0.001),and then to 11.85±2.48 mmHg at 3750 m ASL(P=0.036 compared to IOP at 2261 m ASL)and partially recovered to 13.47±2.57 mmHg upon return to 44 m.IOP in the LA group at HA and VHA was comparable to that in the local residents(12.2±2.4 mmHg for HA,11.5±1.8 mmHg for VHA).IOP was positively associated with SpO2 while negatively associated with pulse rate.CONCLUSION IOP in the LA group gradually reduced as altitude elevated in stages and became comparable to IOP in local residents.Hypoxia may be associated with IOP,which deserves further study.

Forest structure and carbon dynamics of an intact lowland mixed dipterocarp forest in Brunei Darussalam

Tropical forests play a critical role in mitigating climate change because they account for large amount o terrestrial carbon storage and productivity.However,there are many uncertainties associated with the estimation o carbon dynamics.We estimated forest structure and carbon dynamics along a slope（17.3°–42.8°）and to assess the relations between forest structures,carbon dynamics,and slopes in an intact lowland mixed dipterocarp forest,in Kuala Belalong,Brunei Darussalam.Living biomass,basa area,stand density,crown properties,and tree family composition were measured for forest structure.Growth rate,litter production,and litter decomposition rates were also measured for carbon dynamics.The crown form index and the crown position index were used to assess crown properties,which we categorized into five stages,from very poor to perfect.The living biomass,basal area and stand density were 261.5–940.7 Mg ha-1,43.6–63.6 m2ha-1and 6,675–8400 tree ha-1,respectively.The average crown form and position index were 4,which means that the crown are mostly symmetrical and sufficiently exposed for photosynthesis.The mean biomass growth rate,litter production,litter decomposition rate were estimated as11.9,11.6 Mg ha-1a-1,and 7.2 g a-1,respectively.Biomass growth rate was significantly correlated with living biomass,basal area,and crown form.Crown form appeared to strongly influence living biomass,basal area and biomass growth rate in terms of light acquisition.However,basal area,stand density,crown properties,and biomass growth rate did not vary by slope or tree family composition.The results indicate that carbon accumulation by tree growth in an intact lowland mixed dipterocarp forest depends on crown properties.Absence of any effect of tree family composition on carbon accumulation suggests that the main driver of biomass accumulation in old-growth forests of Borneo is not species-specific characteristics of tree species.