基金This w ork is supported by the National N atural Science Foundation of China N o 30 170 0 76 Chinese Academ y of Sciences and the Changbai Mountain Station for Forest Ecosystem Research.
基金General Project of Scientific Research of Educational Commission of Liaoning Province Under Grant(L2015359)Mentoring Program of Natural Science Foundation of Liaoning Province Under Grant(20170540642)
基金supported by Knowledge Innovation Programs of Chinese Academy of Sciences ( KZCX2-YW -BR-16)National Basic Research Program of China ( 973 Program,2010CB951304-4)the National Natural Science Foundation of China ( 40901036,41171047)