This paper focuses on the indicators of soil and litter health, disturbance, and landscape heterogeneity as a tool for prediction of ecosystem sustainability in the northern forests of Iran. The study area was divided...This paper focuses on the indicators of soil and litter health, disturbance, and landscape heterogeneity as a tool for prediction of ecosystem sustainability in the northern forests of Iran. The study area was divided into spatial homogenous sites using slope, aspect, and soil humidity classes. Then a range of sites along the disturbance gradient was selected for sampling. Chemical and physical indicators of soil and litter health were measured at random points within these sites. Structural equation modeling(SEM) was applied to link six constructs of landscape heterogeneity, three constructs of disturbance(harvest, livestock, and human accessibility), and soil and litter health. The results showed that with decreasing accessibility, the total N and organic matter content of soil increased and effective bulk density decreased. Harvesting activities increased soil organic matter. Therefore, it is concluded that disturbances through harvesting and accessibility inversely affect the soil health. Unexpectedly, it was found that the litter total C and C:N ratio improved with an increase in the harvest and accessibility disturbances, whereas litter bulk density decreased. Investigation of tree composition revealed that in the climax communities, which are normally affected more by harvesting activities, some species like Fagus orientalis Lipsky with low decomposition rate are dominant. The research results showed that changes in disturbance intensity are reflected in litter and soil indicators, whereas the SEM indicated that landscape heterogeneity has a moderator effect on the disturbance to both litter and soil paths.展开更多
We studied the long-term impacts of natural burning on herbaceous species diversity 37 years after a fire occurred in the Roudbar forests of Guilan Province, northern Iran. Numerous studies have examined short-term ch...We studied the long-term impacts of natural burning on herbaceous species diversity 37 years after a fire occurred in the Roudbar forests of Guilan Province, northern Iran. Numerous studies have examined short-term changes in understory vegetation following wildfire; however, very few long-term studies are available or changes inferred from retrospective studies based on chro- nosequences. For this study, 170 ha of forest (85 ha, burned areas; 85 ha, unburned areas) were surveyed. Because the 1000 m2 plots were too large for detailed measurements of herbaceous species, we determined a sub-sample size according to the Whittaker's nested plot sampling protocol and minimal areas method. Hence, sub-plots of 32 m2 were used for herbaceous species measurements, which consisted of percent cover of each species based on the Domin criterion. We measured plant diversity (Shannon-Wiener index), spe- cies richness (Margalef's index), and evenness (Smith-Wilson index). Mean percent cover, together with diversity, richness and even- ness, increased markedly in burned areas compared to unburned controls. This suggests that the biodiversity of these forests could be restored within 37 years after fire. However, the abundance of invasive species such as Rubus fruticosus and Bromus benekenii increased significantly in burned areas, but these could be controlled by relevant silvicultural operations.展开更多
The Grass Snake, Natrix natrix, is one of the non-venomous oviparous snakes of the family Colubridae, found in Mazandaran, a northern province of Iran. In order to study the spermatogenic cycle of this species in the ...The Grass Snake, Natrix natrix, is one of the non-venomous oviparous snakes of the family Colubridae, found in Mazandaran, a northern province of Iran. In order to study the spermatogenic cycle of this species in the area, sampling was done from April to November, and twenty-six male snakes were collected by hand during daylight hours. Testes were extracted from the specimens, and were fixed and prepared for histological examination. The results indicated that the spermatogenesis started in early June, November. The mean body length of the specimens was reached its height in August and September, and ended in measured 636.21 mm, and the mean volume of testes was 524.54 rnm3 and 569.06 mm3 for the right and left ones, respectively, and the mean weight of the testes was 0.051 g Spermatogenesis in this species is aestival and sperms are stored for use in the year following production.展开更多
It has become necessary to develop stands of fast growing endemic species in the Caspian forests because of their reduction in area and the domestic demand of timber processing industries in Iran. Acer velutinum, an e...It has become necessary to develop stands of fast growing endemic species in the Caspian forests because of their reduction in area and the domestic demand of timber processing industries in Iran. Acer velutinum, an endemic species in Iran, is often planted in monocultures or in a mixture with other endemic species. Our study was conducted on A. velutinum planted in Berenjestanak(control plots) and Chaibagh(thinned plots) in northern Iran in 1991. A randomized sampling method with circular sampling plots of 100 m2was used to measure diameter and height of trees. The results show that the diameter of trees in thinned stands was 19.98 cm and in the control stands 13.84 cm, while the mean heights were 18.33 and 17.46 m, respectively. The mean basal area in thinned stands was 30.74 m2and in the control 15.70 m2per ha. The results of Student’s t tests indicate that there are significant differences at a probability level of 99.9% between the two variously treated stands in diameter(d), height(h), basal area, volume growth and the stability coefficient. We conclude that tending operations with suitable intensity in different time periods are recommended to obtain stable stands.展开更多
High resolution sampling across the Cretaceous/Paleogene boundary (K/Pg) at the Galanderud section in northern Iran provides the most expanded and continuous section for us to consider biostratigraphy and the mass ext...High resolution sampling across the Cretaceous/Paleogene boundary (K/Pg) at the Galanderud section in northern Iran provides the most expanded and continuous section for us to consider biostratigraphy and the mass extinction pattern of Cretaceous planktic foraminifera. Based on planktic foraminifera, four biozones and five sub-biozones have been identified across the K/Pg boundary. These biozones include the Abathomphalus mayaroensis Biozone (Plummerita hantkeninoides subbiozone), the Guembelitria cretacea Biozone (including two sub-biozones: the Hedbergella holmdelensis and the Parvularugoglobigerina longiapertura), the Parvularugoglobigerina eugubina Biozone (including two subbiozones: the Parvularugoglobigerina Sabina and the Eoglobigerina simplicissima) and finely the Parasubbotina pseudobulloides Biozone. Planktic foraminiferal extinction occurred over a brief period, with 3% of the species disappearing in the late Maastrichtian, and 72% of the species becaming extinct at the K/Pg boundary. About 25% of the species survived into the early Danian. Extinction of 73% of the species at the K/Pg boundary is very compatible with the effect of a large asteroid impact.展开更多
Bark biomass as an energy source has a high economic value.Bark content variations and production helps recognize the potential of this bioenergy source spatially before harvesting.The percentage of fresh and dry bark...Bark biomass as an energy source has a high economic value.Bark content variations and production helps recognize the potential of this bioenergy source spatially before harvesting.The percentage of fresh and dry bark in Populus deltoides grown under a monoculture system was examined in the temperate region of northern Iran.Diameter at breast height(DBH)and total height data were analyzed based on an initial inventory.Ten sample trees were felled,separated into 2 m-segments,and weighted in the field.A 5-cm-thick disc from each segment was extracted for determining fresh and dry bark percentages.These were statistically significantly different in disc diameter classes and decreased with increasing disc diameters.Bark percentage of the disc classes ranged from 21.8 to 24.4%in small-sized diameters to 8.1-9.3%in large-sized diameters.The differences between fresh and dry bark percentages depended on water content variations.Allometric power equations were fitted to data of fresh and dry bark percentages and disc diameters as well as DBH.The values of R~2 ranged from 0.89 to 0.90.In addition,allometric power equations provided the best fits for relationships between total stem dry biomass,dry bark biomass,and DBH,R~2=0.986 and 0.979 for the total stem dry biomass and stem dry bark biomass,respectively.The allometric models can be used to estimate bark percentage and bark production of P.deltoides in segments and for the whole stem for a wide range of segment diameters(8-44 cm)and DBH(15-45 cm).展开更多
文摘This paper focuses on the indicators of soil and litter health, disturbance, and landscape heterogeneity as a tool for prediction of ecosystem sustainability in the northern forests of Iran. The study area was divided into spatial homogenous sites using slope, aspect, and soil humidity classes. Then a range of sites along the disturbance gradient was selected for sampling. Chemical and physical indicators of soil and litter health were measured at random points within these sites. Structural equation modeling(SEM) was applied to link six constructs of landscape heterogeneity, three constructs of disturbance(harvest, livestock, and human accessibility), and soil and litter health. The results showed that with decreasing accessibility, the total N and organic matter content of soil increased and effective bulk density decreased. Harvesting activities increased soil organic matter. Therefore, it is concluded that disturbances through harvesting and accessibility inversely affect the soil health. Unexpectedly, it was found that the litter total C and C:N ratio improved with an increase in the harvest and accessibility disturbances, whereas litter bulk density decreased. Investigation of tree composition revealed that in the climax communities, which are normally affected more by harvesting activities, some species like Fagus orientalis Lipsky with low decomposition rate are dominant. The research results showed that changes in disturbance intensity are reflected in litter and soil indicators, whereas the SEM indicated that landscape heterogeneity has a moderator effect on the disturbance to both litter and soil paths.
文摘We studied the long-term impacts of natural burning on herbaceous species diversity 37 years after a fire occurred in the Roudbar forests of Guilan Province, northern Iran. Numerous studies have examined short-term changes in understory vegetation following wildfire; however, very few long-term studies are available or changes inferred from retrospective studies based on chro- nosequences. For this study, 170 ha of forest (85 ha, burned areas; 85 ha, unburned areas) were surveyed. Because the 1000 m2 plots were too large for detailed measurements of herbaceous species, we determined a sub-sample size according to the Whittaker's nested plot sampling protocol and minimal areas method. Hence, sub-plots of 32 m2 were used for herbaceous species measurements, which consisted of percent cover of each species based on the Domin criterion. We measured plant diversity (Shannon-Wiener index), spe- cies richness (Margalef's index), and evenness (Smith-Wilson index). Mean percent cover, together with diversity, richness and even- ness, increased markedly in burned areas compared to unburned controls. This suggests that the biodiversity of these forests could be restored within 37 years after fire. However, the abundance of invasive species such as Rubus fruticosus and Bromus benekenii increased significantly in burned areas, but these could be controlled by relevant silvicultural operations.
文摘The Grass Snake, Natrix natrix, is one of the non-venomous oviparous snakes of the family Colubridae, found in Mazandaran, a northern province of Iran. In order to study the spermatogenic cycle of this species in the area, sampling was done from April to November, and twenty-six male snakes were collected by hand during daylight hours. Testes were extracted from the specimens, and were fixed and prepared for histological examination. The results indicated that the spermatogenesis started in early June, November. The mean body length of the specimens was reached its height in August and September, and ended in measured 636.21 mm, and the mean volume of testes was 524.54 rnm3 and 569.06 mm3 for the right and left ones, respectively, and the mean weight of the testes was 0.051 g Spermatogenesis in this species is aestival and sperms are stored for use in the year following production.
文摘It has become necessary to develop stands of fast growing endemic species in the Caspian forests because of their reduction in area and the domestic demand of timber processing industries in Iran. Acer velutinum, an endemic species in Iran, is often planted in monocultures or in a mixture with other endemic species. Our study was conducted on A. velutinum planted in Berenjestanak(control plots) and Chaibagh(thinned plots) in northern Iran in 1991. A randomized sampling method with circular sampling plots of 100 m2was used to measure diameter and height of trees. The results show that the diameter of trees in thinned stands was 19.98 cm and in the control stands 13.84 cm, while the mean heights were 18.33 and 17.46 m, respectively. The mean basal area in thinned stands was 30.74 m2and in the control 15.70 m2per ha. The results of Student’s t tests indicate that there are significant differences at a probability level of 99.9% between the two variously treated stands in diameter(d), height(h), basal area, volume growth and the stability coefficient. We conclude that tending operations with suitable intensity in different time periods are recommended to obtain stable stands.
文摘High resolution sampling across the Cretaceous/Paleogene boundary (K/Pg) at the Galanderud section in northern Iran provides the most expanded and continuous section for us to consider biostratigraphy and the mass extinction pattern of Cretaceous planktic foraminifera. Based on planktic foraminifera, four biozones and five sub-biozones have been identified across the K/Pg boundary. These biozones include the Abathomphalus mayaroensis Biozone (Plummerita hantkeninoides subbiozone), the Guembelitria cretacea Biozone (including two sub-biozones: the Hedbergella holmdelensis and the Parvularugoglobigerina longiapertura), the Parvularugoglobigerina eugubina Biozone (including two subbiozones: the Parvularugoglobigerina Sabina and the Eoglobigerina simplicissima) and finely the Parasubbotina pseudobulloides Biozone. Planktic foraminiferal extinction occurred over a brief period, with 3% of the species disappearing in the late Maastrichtian, and 72% of the species becaming extinct at the K/Pg boundary. About 25% of the species survived into the early Danian. Extinction of 73% of the species at the K/Pg boundary is very compatible with the effect of a large asteroid impact.
文摘Bark biomass as an energy source has a high economic value.Bark content variations and production helps recognize the potential of this bioenergy source spatially before harvesting.The percentage of fresh and dry bark in Populus deltoides grown under a monoculture system was examined in the temperate region of northern Iran.Diameter at breast height(DBH)and total height data were analyzed based on an initial inventory.Ten sample trees were felled,separated into 2 m-segments,and weighted in the field.A 5-cm-thick disc from each segment was extracted for determining fresh and dry bark percentages.These were statistically significantly different in disc diameter classes and decreased with increasing disc diameters.Bark percentage of the disc classes ranged from 21.8 to 24.4%in small-sized diameters to 8.1-9.3%in large-sized diameters.The differences between fresh and dry bark percentages depended on water content variations.Allometric power equations were fitted to data of fresh and dry bark percentages and disc diameters as well as DBH.The values of R~2 ranged from 0.89 to 0.90.In addition,allometric power equations provided the best fits for relationships between total stem dry biomass,dry bark biomass,and DBH,R~2=0.986 and 0.979 for the total stem dry biomass and stem dry bark biomass,respectively.The allometric models can be used to estimate bark percentage and bark production of P.deltoides in segments and for the whole stem for a wide range of segment diameters(8-44 cm)and DBH(15-45 cm).