Background:Chinese pine(Pinus tabuliformis Carr.)is one of the major afforestation species in northern China and plays a key role in restoring forest ecosystems and preserving soil and water.However,most Chinese pine ...Background:Chinese pine(Pinus tabuliformis Carr.)is one of the major afforestation species in northern China and plays a key role in restoring forest ecosystems and preserving soil and water.However,most Chinese pine plantations are experiencing ecological problems such as the low diversity of understory plants and difficulty in natural regeneration.Thinning has been widely used to maintain and improve a variety of forest ecosystem services from plantations.To date,however,few studies have been conducted to systematically determine the effects of thinning on understory plant diversity and the regeneration of Chinese pine in plantations.Methods:We conducted a literature search,and selected 22 publications covering a total of 83 treatments related to thinning effects on the species richness of understory plants and 15 publications covering a total of 43 treatments related to thinning effects on the regeneration of Chinese pine,in tree plantations of northern China.The data from the literature were synthesized and evaluated with meta-analysis approach to determine the treatment effects.Results:Compared with the control stands,thinning increased the species richness of shrubs and herbs by an average of 25.3%and 26.5%,respectively.While the varying thinning intensities all had significantly positive effects on the species richness of understory plants,only moderate thinning(30%–50%)had a positive effect on the density of regenerating seedlings and saplings of Chinese pine(60.2%).The species richness of understory plants was greatest after 14 years of thinning with an increase of 36.3%,whereas the density of regenerating Chinese pine seedlings and saplings reached a maximum after≥11 years of thinning with an increase of 76.5%,compared to that of the unthinned stands.Thinning in the half-mature plantations had the greatest effects on the understory shrub richness(44.1%)and the density of regenerating Chinese pine seedlings and saplings(86.5%).Both single and multiple thinning were found to significantly promote the species richness of understory plants and the density of regenerating Chinese pine seedlings and saplings,and the positive effects of thinning were greater in areas with a humidity index(HI)<30 than in areas with an HI≥30.In general,age group,planting density and recovery time were prominent factors affecting the species richness of understory plants,whereas the slope,HI and recovery time were the dominant controls of the density of regenerating Chinese pine seedlings and saplings,indicating differential effects of thinning on the species richness of understory plants and the regeneration capacity of Chinese pine in plantations.Conclusion:Thinning appears to be a feasible management measure to improve the understory plant diversity and regeneration capacity of Chinese pine in plantations.We postulate that moderate thinning in half-mature forest stands with an HI<30 can help effectively promote the species diversity of understory plants and the natural regeneration of Chinese pine,thereby maintaining a more resilient stand structure and the development of Chinese pine plantations.展开更多
In 2014,a 40-ha undisturbed plot was established in the Beijing Songshan Nature Reserve.The spatial distribution and spatial associations of five dominant tree species of different height classes were examined using s...In 2014,a 40-ha undisturbed plot was established in the Beijing Songshan Nature Reserve.The spatial distribution and spatial associations of five dominant tree species of different height classes were examined using spatial point pattern analysis and the effects of these species on local community diversity assemblages were examined using the ISAR method.The dominant species were characterized by an aggregated distribution at spatial scales of 0–50 m.At smaller spatial scales,trees in larger height classes had significant positive and negative effects on the diversity of the lower height classes.However,at larger spatial scales this effect was diminished.At small scales(0–10 m),accumulator species had positive effects on species diversity,maintained an over-representative proportion of diversity in their proximity,and supported the ecological niche theory.At the larger scale(10–50 m),neutral species were dominant and had positive effects on species diversity,though repeller species,which had negative effects on species diversity,also contributed to diversity.Neutral and accumulator species together determined local species diversity,but the relative importance of the two was closely related to spatial scale.A combination of the ecological niche theory and neutral processes together determines species coexistence and biodiversity of an undisturbed pine forest.展开更多
基金supported by Natural Science Basic Research Program of Shaanxi(2021JQ-155)National Natural Science Foundation of China(32101511)Chinese Universities Scientific Fund(2452020137 and 2452021073)。
文摘Background:Chinese pine(Pinus tabuliformis Carr.)is one of the major afforestation species in northern China and plays a key role in restoring forest ecosystems and preserving soil and water.However,most Chinese pine plantations are experiencing ecological problems such as the low diversity of understory plants and difficulty in natural regeneration.Thinning has been widely used to maintain and improve a variety of forest ecosystem services from plantations.To date,however,few studies have been conducted to systematically determine the effects of thinning on understory plant diversity and the regeneration of Chinese pine in plantations.Methods:We conducted a literature search,and selected 22 publications covering a total of 83 treatments related to thinning effects on the species richness of understory plants and 15 publications covering a total of 43 treatments related to thinning effects on the regeneration of Chinese pine,in tree plantations of northern China.The data from the literature were synthesized and evaluated with meta-analysis approach to determine the treatment effects.Results:Compared with the control stands,thinning increased the species richness of shrubs and herbs by an average of 25.3%and 26.5%,respectively.While the varying thinning intensities all had significantly positive effects on the species richness of understory plants,only moderate thinning(30%–50%)had a positive effect on the density of regenerating seedlings and saplings of Chinese pine(60.2%).The species richness of understory plants was greatest after 14 years of thinning with an increase of 36.3%,whereas the density of regenerating Chinese pine seedlings and saplings reached a maximum after≥11 years of thinning with an increase of 76.5%,compared to that of the unthinned stands.Thinning in the half-mature plantations had the greatest effects on the understory shrub richness(44.1%)and the density of regenerating Chinese pine seedlings and saplings(86.5%).Both single and multiple thinning were found to significantly promote the species richness of understory plants and the density of regenerating Chinese pine seedlings and saplings,and the positive effects of thinning were greater in areas with a humidity index(HI)<30 than in areas with an HI≥30.In general,age group,planting density and recovery time were prominent factors affecting the species richness of understory plants,whereas the slope,HI and recovery time were the dominant controls of the density of regenerating Chinese pine seedlings and saplings,indicating differential effects of thinning on the species richness of understory plants and the regeneration capacity of Chinese pine in plantations.Conclusion:Thinning appears to be a feasible management measure to improve the understory plant diversity and regeneration capacity of Chinese pine in plantations.We postulate that moderate thinning in half-mature forest stands with an HI<30 can help effectively promote the species diversity of understory plants and the natural regeneration of Chinese pine,thereby maintaining a more resilient stand structure and the development of Chinese pine plantations.
基金supported by the Beijing Common Construction Project Research and demonstrations on the regression technique for minimum populations of wild plants(2016YFC0503106)
文摘In 2014,a 40-ha undisturbed plot was established in the Beijing Songshan Nature Reserve.The spatial distribution and spatial associations of five dominant tree species of different height classes were examined using spatial point pattern analysis and the effects of these species on local community diversity assemblages were examined using the ISAR method.The dominant species were characterized by an aggregated distribution at spatial scales of 0–50 m.At smaller spatial scales,trees in larger height classes had significant positive and negative effects on the diversity of the lower height classes.However,at larger spatial scales this effect was diminished.At small scales(0–10 m),accumulator species had positive effects on species diversity,maintained an over-representative proportion of diversity in their proximity,and supported the ecological niche theory.At the larger scale(10–50 m),neutral species were dominant and had positive effects on species diversity,though repeller species,which had negative effects on species diversity,also contributed to diversity.Neutral and accumulator species together determined local species diversity,but the relative importance of the two was closely related to spatial scale.A combination of the ecological niche theory and neutral processes together determines species coexistence and biodiversity of an undisturbed pine forest.
