Vertical forest structure is closely linked to multiple ecosystem characteristics,such as biodiversity,habitat,and productivity.Mixing tree species in planted forests has the potential to create diverse vertical fores...Vertical forest structure is closely linked to multiple ecosystem characteristics,such as biodiversity,habitat,and productivity.Mixing tree species in planted forests has the potential to create diverse vertical forest structures due to the different physiological and morphological traits of the composing tree species.However,the relative importance of species richness,species identity and species interactions for the variation in vertical forest structure remains unclear,mainly because traditional forest inventories do not observe vertical stand structure in detail.Terrestrial laser scanning(TLS),however,allows to study vertical forest structure in an unprecedented way.Therefore,we used TLS single scan data from 126 plots across three experimental planted forests of a largescale tree diversity experiment in Belgium to study the drivers of vertical forest structure.These plots were 9–11years old young pure and mixed forests,characterized by four levels of tree species richness ranging from monocultures to four-species mixtures,across twenty composition levels.We generated vertical plant profiles from the TLS data and derived six stand structural variables.Linear mixed models were used to test the effect of species richness on structural variables.Employing a hierarchical diversity interaction modelling framework,we further assessed species identity effect and various species interaction effects on the six stand structural variables.Our results showed that species richness did not significantly influence most of the stand structure variables,except for canopy height and foliage height diversity.Species identity on the other hand exhibited a significant impact on vertical forest structure across all sites.Species interaction effects were observed to be site-dependent due to varying site conditions and species pools,and rapidly growing tree species tend to dominate these interactions.Overall,our results highlighted the importance of considering both species identity and interaction effects in choosing suitable species combinations for forest management practices aimed at enhancing vertical forest structure.展开更多
Cloud is essential in the atmosphere, condensing water vapor and generating strong convective or large-scale persistent precipitation. In this work, the relationships between cloud vertical macro- or microphysical pro...Cloud is essential in the atmosphere, condensing water vapor and generating strong convective or large-scale persistent precipitation. In this work, the relationships between cloud vertical macro- or microphysical properties, radiative heating rate, and precipitation for convective and stratiform clouds in boreal summer over the Tibetan Plateau (TP) are analyzed and compared with its neighboring land and tropical oceans based on CloudSat/CALIPSO satellite measurements and TRMM precipitation data. The precipitation intensity caused by convective clouds is twofold stronger than that by stratiform clouds. The vertical macrophysics of both cloud types show similar features over the TP, with the region weakening the precipitation intensity and compressing the cloud vertical expansion and variation in cloud top height, but having an uplift effect on the average cloud top height. The vertical microphysics of both cloud types under conditions of no rain over the TP are characterized by lower-level ice water, ice particles with a relatively larger range of sizes, and a relatively lower occurrence of denser ice particles. The features are similar to other regions when precipitation enhances, but convective clouds gather denser and larger ice particles than stratiform clouds over the TP. The atmospheric shortwave (longwave) heating (cooling) rate strengthens with increased precipitation for both cloud types. The longwave cooling layer is thicker when the rainfall rate is less than 100 mm d?1, but the net heating layer is typically compressed for the profiles of both cloud types over the TP. This study provides insights into the associations between clouds and precipitation, and an observational basis for improving the simulation of convective and stratiform clouds over the TP in climate models.展开更多
Knowledge of the vertical distribution of aerosols in the free troposphere is important for estimating their impact on climate. In this study, direct observations of the vertical distribution of aerosols in the free t...Knowledge of the vertical distribution of aerosols in the free troposphere is important for estimating their impact on climate. In this study, direct observations of the vertical distribution of aerosols in the free troposphere are made using surface Micro-Pulse Lidar (MPL) measurements. The MPL measurements were made at the Loess Plateau (35.95°N, 104.1°E), which is near the major dust source regions of the Taklimakan and Gobi deserts. The vertical distribution of the MPL backscattering suggested that nondust aerosols floated from ground level to an altitude of approximately 9 km around the source regions. Early morning hours are characterized by a shallow aerosol layer of a few hundred meters thick. As the day progresses, strong convective eddies transport the aero- sols vertically to more than 1500 m.展开更多
ABSTRACT The abilities of BCC-AGCM2.1 and BCC_AGCM2.2 to simulate the annual-mean cloud vertical structure (CVS) were evaluated through comparison with GCM-Oriented CALIPSO Cloud Product (CALIPSO-GOCCP) data. BCC...ABSTRACT The abilities of BCC-AGCM2.1 and BCC_AGCM2.2 to simulate the annual-mean cloud vertical structure (CVS) were evaluated through comparison with GCM-Oriented CALIPSO Cloud Product (CALIPSO-GOCCP) data. BCC-AGCM2.2 has a dynamical core and physical processes that are consistent with BCC-AGCM2.1, but has a higher horizontal resolution. Results showed that both BCC-AGCM versions underestimated the global-mean total cloud cover (TCC), middle cloud cover (MCC) and low cloud cover (LCC), and that BCC_AGCM2.2 underestimated the global-mean high cloud cover (HCC). The global-mean cloud cover shows a systematic decrease from BCCA-GCM2.1 to BCC_AGCM2.2, especially for HCC. Geographically, HCC is significantly overestimated in the tropics, particularly by BCC_AGCM2,1, while LCC is generally overestimated over extra-tropical lands, but significantly underestimated over most of the oceans, especially for subtropical marine stratocumulus clouds. The leading EOF modes of CVS were extracted. The BCC_AGCMs perform well in reproducing EOF1, but with a larger variance explained. The two models also capture the basic features of EOF3, except an obvious deficiency in eigen- vector peaks. EOF2 has the largest simulation biases in both position and strength of eigenvector peaks. Furthermore, we investigated the effects of CVS on relative shortwave and longwave cloud radiative forcing (RSCRF and RLCRF). Both BCC_AGCM versions successfully reproduce the sign of regression coefficients, except for RLCRF in PC1. However, the RSCRF relative contributions from PC1 and PC2 are overestimated, while the relative contribution from PC3 is underes timated in both BCC_AGCM versions. The RLCRF relative contribution is underestimated for PC2 and overestimated for PC3.展开更多
A comprehensive protective structure with rigidity and flexibility was put forward and designed in view of the quality and safety problems for the double vertical explosive welding of large titanium/steel cladding pla...A comprehensive protective structure with rigidity and flexibility was put forward and designed in view of the quality and safety problems for the double vertical explosive welding of large titanium/steel cladding plate.The movement speed and displacement of the protective structure was calculated by establishing its physics model.The dynamics and stabilization properties were analyzed,and the protective structure parameters were optimized and devised.The comprehensive protective structure,which is composed of rigidity unit and flexibility wall,can bear the impact of detonation wave and the high-speed movement of the cladding plate.There are no damage and deformation in the protective structure and the cladding plate.The protective structure can be used many times.The bonding rate of the Ti/steel plate obtained was nearly 100%,and there is no deformation,surface cracks,and big wave and micro-defects.Therefore,the protective problems of the double vertical explosive welding can be solved effectively by the protective structure.展开更多
The solidified structure of the thin-walled and complicated Ti-6AI-4V castings produced by the vertical centrifugal casting process was studied in the present work. The results show that the wall thickness of the sect...The solidified structure of the thin-walled and complicated Ti-6AI-4V castings produced by the vertical centrifugal casting process was studied in the present work. The results show that the wall thickness of the section is featured with homogeneously distributed fine equiaxial grains, compared with the microstructure of the thick-walled section. The grain size of the castings has a tendency to decrease gradually with the increasing of the centrifugal radius. The inter-lamellar space in thick-walled casting parts is bigger than that of the thin-walled parts, and the profile of inter-lamellar space is not susceptible to the centrifugal radius.展开更多
In the ice-covered oceanic region,the collision between sea ice and offshore structures will occur,causing the crushing failure of ice and the vibration of structures.The vibration can result in fatigue damage of stru...In the ice-covered oceanic region,the collision between sea ice and offshore structures will occur,causing the crushing failure of ice and the vibration of structures.The vibration can result in fatigue damage of structure and even endanger the crews’health.It is no doubt that this ice-structure interaction has been noted with great interest by the academic community for a long time and numerous studies have been done through theoretical analysis,experimental statistics and numerical simulation.In this paper,the bond-based Peridynamics method is applied to simulate the interaction between sea ice and wide vertical structures,where sea ice is modeled as elastic-plastic material,with a certain yield condition and failure criterion.Oscillation equation of single-degree-of-freedom is considered to investigate the vibration features of the structure during the interaction process.The damage of ice,ice forces and vibration responses of structure in the duration are obtained through numerical simulation.A parametric investigation is undertaken to identify the key parameters,such as ice thickness,the diameter of structure and relative velocity that trigger the ice crushing,ice forces and vibration responses of the structure.Results indicate that all three parameters have a positive correlation with the overall level of ice force and vibration displacement.Besides,a velocity coefficient is proposed to predict the vibration displacement based on its relation with ice speed.展开更多
This study investigates the vertical structure of variability in the cross-equatorial flows(CEFs)over the Maritime Continent and Indian Ocean in boreal summer,based on three reanalysis datasets:ERA-Interim,JRA-55 and ...This study investigates the vertical structure of variability in the cross-equatorial flows(CEFs)over the Maritime Continent and Indian Ocean in boreal summer,based on three reanalysis datasets:ERA-Interim,JRA-55 and NCEP-2.The results show a high consistency in the interannual variability among the reanalysis datasets,especially between ERAInterim and JRA-55,while great uncertainty exists in the decadal or long-term changes of CEFs.Further analyses on the interannual variability in CEFs indicate that there is a significant negative relationship between the upper-and lower-level CEFs over the Maritime Continent—that is,the northerlies at the upper level and southerlies at the lower level are both enhanced or weakened.This seesaw pattern is also significantly related to the CEFs over the Indian Ocean at the upper level and lower level(i.e.,the Somali jet).This close relationship between the upper-and lower-level CEFs and between the Maritime Continent and Indian Ocean is manifested as the leading mode of equatorial meridional winds in the vertical-zonal section over the Maritime Continent and Indian Ocean.Finally,it is found that ENSO is closely related to the vertical leading mode,and to all the CEFs at the upper and lower levels over the Maritime Continent and Indian Ocean.展开更多
With the method of separation of variables and the eigenfunction expansion employed, an analytical solution is presented for the radiation and diffraction of a rectangular structure with an opening near a vertical wal...With the method of separation of variables and the eigenfunction expansion employed, an analytical solution is presented for the radiation and diffraction of a rectangular structure with an opening near a vertical wall in oblique seas, in which the unknown coefficients are determined by the boundary conditions and matching requirement on the interface. The effects of the width of the opening and the angle of incidence on the hydrodynamic characteristics of a rectangular structure with an opening near a vertical wall are mainly studied. The comparisons of the calculation results with wall-present and with wall-absent are also made. The results indicate that the variation trends of the heave added mass and excitation force with wall-present are almost the same as those with wall-absent, and that the peak values in the former case are obviously larger than those in the latter due to the reflection of the vertical wall.展开更多
A new type of vibration structure (i.e. supporting system, called swing frame cus- tomarily) of vertical dynamic balancing machine has been designed, which is based on an analysis for the swing frame of a traditiona...A new type of vibration structure (i.e. supporting system, called swing frame cus- tomarily) of vertical dynamic balancing machine has been designed, which is based on an analysis for the swing frame of a traditional double-plane vertical dynamic balancing machine. The static unbalance and couple unbalance can be e?ectively separated by using the new dynamic balancing machine with the new swing frame. By building the dynamics model, the advantages of the new structure are discussed in detail. The modal and harmonic response are analyzed by using the ANSYS7.0. By comparing the ?nite element modal analysis with the experimental modal analy- sis, the natural frequencies and vibration modes are found. There are many spring boards in the new swing frame. Their sti?nesses are di?erent and assorted with each other. Furthermore, there are three sensors on the measuring points. Therefore, the new dynamic balancing machine can measure static unbalance and coupling unbalance directly, and the interaction between them is faint. The result shows that the new vertical dynamic balancing machine is suitable for inertial measurement of ?ying objects, and can overcome the shortcomings of traditional double-plane vertical dynamic balancing machines, which the e?ect of plane-separation is inferior. The vertical dynamic balancing machine with the new vibration structure can ?nd wide application in the future. The modelling and analysis of the new vibration structure will provide theoretical basis and practical experience for designing new-type vertical dynamic balancing machines.展开更多
Empirical Orthogonal Function (EOF) analysis is used in this study to generate main eigenvector fields of historical temperature for the China Seas (here referring to Chinese marine territories) and adjacent water...Empirical Orthogonal Function (EOF) analysis is used in this study to generate main eigenvector fields of historical temperature for the China Seas (here referring to Chinese marine territories) and adjacent waters from 1930 to 2002 (510 143 profiles). A good temperature profile is reconstructed based on several subsurface in situ temperature observations and the thermocline was estimated using the model. The results show that: 1) For the study area, the former four principal components can explain 95% of the overall variance, and the vertical distribution of temperature is most stable using the in situ temperature observations near the surface. 2) The model verifications based on the observed CTD data from the East China Sea (ECS), South China Sea (SCS) and the areas around Taiwan Island show that the reconstructed profiles have high correlation with the observed ones with the confidence level 〉95%, especially to describe the characteristics of the thermocline well. The average errors between the reconstructed and observed profiles in these three areas are 0.69℃, 0.52℃ and 1.18℃ respectively. It also shows the model RMS error is less than or close to the climatological error. The statistical model can be used to well estimate the temperature profile vertical structure. 3) Comparing the thermocline characteristics between the reconstructed and observed profiles, the results in the ECS show that the average absolute errors are 1.5m, 1.4 m and 0.17~C/m, and the average relative errors are 24.7%, 8.9% and 22.6% for the upper, lower thermocline boundaries and the gradient, respectively. Although the relative errors are obvious, the absolute error is small. In the SCS, the average absolute errors are 4.1 m, 27.7 m and 0.007℃/m, and the average relative errors are 16.1%, 16.8% and 9.5% for the upper, lower thermocline boundaries and the gradient, respectively. The average relative errors are all 〈20%. Although the average absolute error of the lower thermocline boundary is considerable, but contrast to the spatial scale of average depth of the lower thermocline boundary (165 m), the average relative error is small (16.8%). Therefore the model can be used to well estimate the thermocline.展开更多
The sensitive area of targeted observations for short-term(7 d)prediction of vertical thermal structure(VTS)in summer in the Yellow Sea was investigated.We applied the Conditional Nonlinear Optimal Perturbation(CNOP)m...The sensitive area of targeted observations for short-term(7 d)prediction of vertical thermal structure(VTS)in summer in the Yellow Sea was investigated.We applied the Conditional Nonlinear Optimal Perturbation(CNOP)method and an adjoint-free algorithm with the Regional Ocean Modeling System(ROMS).We used vertical integration of CNOP-type temperature errors to locate the sensitive areas,where reduction of initial errors is expected to yield the greatest improvement in VTS prediction for the selected verification area.The identified sensitive areas were northeast−southwest orientated northeast to the verification area,which were possibly related to the southwestward background currents.Then,we performed a series of sensitivity experiments to evaluate the effectiveness of the identified sensitive areas.Results show that initial errors in the identified sensitive areas had the greatest negative effect on VTS prediction in the verification area compared to errors in other areas(e.g.,the verification area and areas to its east and northeast).Moreover,removal of initial errors through deploying simulated observations in the identified sensitive areas led to more refined prediction than correction of initial conditions in the verification area itself.Our results suggest that implementation of targeted observation in the CNOP-based sensitive areas is an effective method to improve short-term prediction of VTS in summer in the Yellow Sea.展开更多
A new type of vibration structure of vertical dynamic balancing machine isdesigned, which is based on the analysis for swing frame of a traditional vertical dynamic balancingmachine. The static unbalance and couple un...A new type of vibration structure of vertical dynamic balancing machine isdesigned, which is based on the analysis for swing frame of a traditional vertical dynamic balancingmachine. The static unbalance and couple unbalance can be separated effectively by using the newmachine with the new swing frame. By building the dynamics model, the advantages of the newstructure are discussed in detail. The modal and harmonic response are analyzed by using theANSYS7.0. By comparing the finite element modal analysis with the experimental modal analysis, thenatural frequencies and vibration modes are found out. There are many spring boards in the new swingframe. Their stiffness is different and assort with each other. Furthermore, there are threesensors on the measurement points. Therefore, the new dynamic balancing machine can measure thestatic unbalance and couple unbalance directly, and the influence between them is faint. The newstructure has the function of belt-strain compensation to improve the measurement precision. Thepractical result indicates that the new vertical dynamic balancing machine is suitable for inertialmeasurement of flying objects, and can overcome the shortcomings of traditional double-planevertical dynamic balancing machines. The vertical dynamic balancing machine with the new vibrationstructure can be widely used in the future applications. The modeling and analysis of the newvibration structure provide theoretic instruction and practical experience for designing new type ofvertical dynamic balancing machines. Based on the design principles such as stiffness-matching,frequency-adjacence and strain-compensation and so on, various new type of vibration structures canbe designed.展开更多
In the coastal ecosystems of the Black Sea,macrophytobenthos and,in particular,the association of Cystoseira crinite,C.barbara,Cladostephus verticillatus,and Corallina mediterranea,with its thick vegetative canopy(VC)...In the coastal ecosystems of the Black Sea,macrophytobenthos and,in particular,the association of Cystoseira crinite,C.barbara,Cladostephus verticillatus,and Corallina mediterranea,with its thick vegetative canopy(VC),is the key contributor to primary production(PP).Though the vertical structure of the canopy,formed by the algal association,is of principal importance to the PP level,this subject has been long-neglected by researchers.The goal of our work was to compare vertical structures of the vegetative canopy of Cystoseira brown algae under diverse hydrodynamical conditions of the Crimean Peninsula coast.Samples were collected using the 50 cm×50 cm counting frame at eight stations positioned in shallow(55-60 cm deep)sites of Sevastopol Bay(Crimean Peninsula).Dry weight biomass of the VC was determined for all algae assemblage and for each algal species individually,per horizontal surface unit,at each height(Z).The study shows that:1)the VC is characterized by unimodal vertical distribution of biomass,with maximum estimate in the lower part,where the biomass increases to 85%of the total biomass;2)a series of single-peaked curves reliably describes the unimodal distribution of the biomass;thalli of different age groups are found along the canopy profile;and 3)algae found in epiphytic synusia prefer inhabiting the upper part of the VC.The role of environmental factors(seawater turbulence and solar radiation)is discussed in reference to the formation of the vertical structure,made up of the associations of the brown algae Cystoseira.展开更多
The algebraic solitary wave and its associated eigenvalue problem in a deep stratified fluid with a free surface and a shallow upper layer were studied. And its vertical structure was examined. An exact solution for t...The algebraic solitary wave and its associated eigenvalue problem in a deep stratified fluid with a free surface and a shallow upper layer were studied. And its vertical structure was examined. An exact solution for the derived 2D Benjamin-Ono equation was obtained, and physical explanation was given with the corresponding dispersion relation. As a special case, the vertical structure of the weakly nonlinear internal wave for the Holmboe density distribution was numerically investigated, and the propagating mechanism of the internal wave was studied by using the ray theory.展开更多
Cloud profiling radar(CPR) onboard Cloud Sat allows for deep penetration into dense clouds/precipitation. In this study, tropical cyclones(TCs) are classified into three stages as developing, mature, and decaying. The...Cloud profiling radar(CPR) onboard Cloud Sat allows for deep penetration into dense clouds/precipitation. In this study, tropical cyclones(TCs) are classified into three stages as developing, mature, and decaying. The circular TC area with the radius of 500 km is divided into five regions. The vertical structure characteristics of 94 Western Pacific TCs at different stages in different regions from June 2006 to February 2014 are statistically quantified using the Cloud Sat tropical cyclone overpass product(the CSTC Product). Contoured frequency by altitude diagrams(CFADs) of radar reflectivity show an arc-like feature and exhibit opposite distributions with a boundary at 5 km. Bright bands are found at this altitude, indicating melting layers. Deep convective(DC) clouds have the largest occurrence probability in the inner region, while Ci clouds occur more frequently in the outer region at 10-15 km. As clouds have the second largest vertical scale after DC clouds. Distributions of Ac, Cu, and Ns clouds at different stages have few distinctions.As the altitude increases, the ice effective radius and the distribution width parameter decrease while the particle number concentration increases. Moist static energy(MSE), cloud thickness(CT), liquid water path(LWP), ice water path(IWP), water vapor(WV), and rain rate(RR) all diminish along the radial direction and are significantly larger at the mature stage. The average value of MSE at the developing stage is larger than that at the decaying stage.展开更多
In this paper, on the basis of the heat conduction equation without consideration of the advection and turbulence effects, one-dimensional model for describing surface sea temperature ( T1), bottom sea temperature ( T...In this paper, on the basis of the heat conduction equation without consideration of the advection and turbulence effects, one-dimensional model for describing surface sea temperature ( T1), bottom sea temperature ( Tt ) and the thickness of the upper homogeneous layer ( h ) is developed in terms of the dimensionless temperature θT and depth η and self-simulation function θT - f(η) of vertical temperature profile by means of historical temperature data.The results of trial prediction with our one-dimensional model on T, Th, h , the thickness and gradient of thermocline are satisfactory to some extent.展开更多
Using the hydroacoustic method with a 200 kHz scientific echo sounding system, the diel vertical migration (DVM) of the sound-scatteringlayer (SSL) in the Yellow Sea Bottom Cold Water (YSBCW) of the southeastern...Using the hydroacoustic method with a 200 kHz scientific echo sounding system, the diel vertical migration (DVM) of the sound-scatteringlayer (SSL) in the Yellow Sea Bottom Cold Water (YSBCW) of the southeastern Yellow Sea was studied in April (spring) and August (summer) of 2010 and 2011. For each survey, 13-27 hours of acoustic data were continuously collected at a stationary station. The acoustic volume scattering strength (Sv) data were analyzed with temperature profile data. In the spring of both 2010 and 2011, the SSL clearly showed the vertical migration throughout the entire water column, moving from the surface layer at night to near the bottom during the day. Conductivity, temperature, and depth data indicated that the entire water column was well mixed with low temperature of about 8℃. However, the SSL showed different patterns in the summers of 2010 and 2011. In the summer of 2010 (≈28℃ at the surface), the SSL migrated to near the bottom during the day, but there were two SSLs above and below the thermocline at depth of 10-30 m at night. In the summer of 2011 (≈20℃ at the surface), the SSL extended throughout the entire water column at night, possibly owing to an abrupt change in sea weather conditions caused by the passage of a Typhoon Muifa over the study area. It was cancluded that the DVM patterns in summer in the YSBCW area may be greatly influenced by a strengthened or weakened thermocline.展开更多
Turbulence in the nocturnal boundary layer(NBL)is still not well characterized,especially over complex underlying surfaces.Herein,gradient tower data and eddy covariance data collected by the Beijing 325-m tower were ...Turbulence in the nocturnal boundary layer(NBL)is still not well characterized,especially over complex underlying surfaces.Herein,gradient tower data and eddy covariance data collected by the Beijing 325-m tower were used to better understand the differentiating characteristics of turbulence regimes and vertical turbulence structure of urban the NBL.As for heights above the urban canopy layer(UCL),the relationship between turbulence velocity scale(VTKE)and wind speed(V)was consistent with the“HOckey-Stick”(HOST)theory proposed for a relatively flat area.Four regimes have been identified according to urban nocturnal stable boundary layer.Regime 1 occurs where local shear plays a leading role for weak turbulence under the constraint that the wind speed V<VT(threshold wind speed).Regime 2 is determined by the existence of strong turbulence that occurs when V>VT and is mainly driven by bulk shear.Regime 3 is identified by the existence of moderate turbulence when upside-down turbulence sporadic bursts occur in the presence of otherwise weak turbulence.Regime 4 is identified as buoyancy turbulence,when V>VT,and the turbulence regime is affected by a combination of local wind shear,bulk shear and buoyancy turbulence.The turbulence activities demonstrated a weak thermal stratification dependency in regime 1,for which within the UCL,the turbulence intensity was strongly affected by local wind shear when V<VT.This study further showed typical examples of different stable boundary layers and the variations between turbulence regimes by analyzing the evolution of wind vectors.Partly because of the influence of large-scale motions,the power spectral density of vertical velocity for upsidedown structure showed an increase at low frequencies.The upside-down structures were also characterized by the highest frequency of the stable stratifications in the higher layer.展开更多
We investigated the vertical variability of toxic potential (i.e. proportions of cells containing microcystin genes) and genotypic structure within different populations of Microcystis aeruginosa that developed in dee...We investigated the vertical variability of toxic potential (i.e. proportions of cells containing microcystin genes) and genotypic structure within different populations of Microcystis aeruginosa that developed in deep artificial reservoirs on the Loire River (France). We demonstrated that a great qualitative vertical heterogeneity could exist within a single bloom of this cyanobacterium in deep lakes. Indeed, we observed important vertical shifts of both toxic potential and genotypic structure, whatever the bloom magnitude. These variations occurred mainly within the euphotic zone and proved to occur independently from abundance vertical shifts. One of the most striking results of this study is that the genotypic structure of a population of M. aeruginosa was more variable between different depths sampled at a single site than between different sites of the same reservoir sampled on top of the water column. In the same way the proportion of potentially toxic cells was sometimes more variable vertically than horizontally. The occurrence of such vertical heterogeneity in three different blooms suggests that this could be a frequent pattern within populations of M. aeruginosa.展开更多
基金Mengxi Wang holds a doctoral scholarship from the China scholarship council(CSC:202003270025)。
文摘Vertical forest structure is closely linked to multiple ecosystem characteristics,such as biodiversity,habitat,and productivity.Mixing tree species in planted forests has the potential to create diverse vertical forest structures due to the different physiological and morphological traits of the composing tree species.However,the relative importance of species richness,species identity and species interactions for the variation in vertical forest structure remains unclear,mainly because traditional forest inventories do not observe vertical stand structure in detail.Terrestrial laser scanning(TLS),however,allows to study vertical forest structure in an unprecedented way.Therefore,we used TLS single scan data from 126 plots across three experimental planted forests of a largescale tree diversity experiment in Belgium to study the drivers of vertical forest structure.These plots were 9–11years old young pure and mixed forests,characterized by four levels of tree species richness ranging from monocultures to four-species mixtures,across twenty composition levels.We generated vertical plant profiles from the TLS data and derived six stand structural variables.Linear mixed models were used to test the effect of species richness on structural variables.Employing a hierarchical diversity interaction modelling framework,we further assessed species identity effect and various species interaction effects on the six stand structural variables.Our results showed that species richness did not significantly influence most of the stand structure variables,except for canopy height and foliage height diversity.Species identity on the other hand exhibited a significant impact on vertical forest structure across all sites.Species interaction effects were observed to be site-dependent due to varying site conditions and species pools,and rapidly growing tree species tend to dominate these interactions.Overall,our results highlighted the importance of considering both species identity and interaction effects in choosing suitable species combinations for forest management practices aimed at enhancing vertical forest structure.
基金jointly supported by the National Natural Science Foundation of China (Grant Nos. 91437219, 91637312 and 91637101)the Key Research Program of Frontier Sciences, Chinese Academy of Sciences (Grant No. QYZDY-SSWDQC018)The CloudSat/CALIPSO data were obtained from the CloudSat Data Processing Center (http://www.cloudsat.cira. colostate.edu/order-data) funded by NASA’s CloudSat project
文摘Cloud is essential in the atmosphere, condensing water vapor and generating strong convective or large-scale persistent precipitation. In this work, the relationships between cloud vertical macro- or microphysical properties, radiative heating rate, and precipitation for convective and stratiform clouds in boreal summer over the Tibetan Plateau (TP) are analyzed and compared with its neighboring land and tropical oceans based on CloudSat/CALIPSO satellite measurements and TRMM precipitation data. The precipitation intensity caused by convective clouds is twofold stronger than that by stratiform clouds. The vertical macrophysics of both cloud types show similar features over the TP, with the region weakening the precipitation intensity and compressing the cloud vertical expansion and variation in cloud top height, but having an uplift effect on the average cloud top height. The vertical microphysics of both cloud types under conditions of no rain over the TP are characterized by lower-level ice water, ice particles with a relatively larger range of sizes, and a relatively lower occurrence of denser ice particles. The features are similar to other regions when precipitation enhances, but convective clouds gather denser and larger ice particles than stratiform clouds over the TP. The atmospheric shortwave (longwave) heating (cooling) rate strengthens with increased precipitation for both cloud types. The longwave cooling layer is thicker when the rainfall rate is less than 100 mm d?1, but the net heating layer is typically compressed for the profiles of both cloud types over the TP. This study provides insights into the associations between clouds and precipitation, and an observational basis for improving the simulation of convective and stratiform clouds over the TP in climate models.
基金supported by the National Natural Science Foundation of China under Grant Nos. 40628005 and 40633017
文摘Knowledge of the vertical distribution of aerosols in the free troposphere is important for estimating their impact on climate. In this study, direct observations of the vertical distribution of aerosols in the free troposphere are made using surface Micro-Pulse Lidar (MPL) measurements. The MPL measurements were made at the Loess Plateau (35.95°N, 104.1°E), which is near the major dust source regions of the Taklimakan and Gobi deserts. The vertical distribution of the MPL backscattering suggested that nondust aerosols floated from ground level to an altitude of approximately 9 km around the source regions. Early morning hours are characterized by a shallow aerosol layer of a few hundred meters thick. As the day progresses, strong convective eddies transport the aero- sols vertically to more than 1500 m.
基金jointly supported by the National Natural Science Foundation of China (Grant Nos.41275077 and 41105054)the National Basic Research Program of China (973 Program:2010CB951902)+1 种基金the China Meteorological Administration (Grant Nos.GYHY201106022 and GYHY201306048)the Sun Yat-sen University "985 Project", Phase 3
文摘ABSTRACT The abilities of BCC-AGCM2.1 and BCC_AGCM2.2 to simulate the annual-mean cloud vertical structure (CVS) were evaluated through comparison with GCM-Oriented CALIPSO Cloud Product (CALIPSO-GOCCP) data. BCC-AGCM2.2 has a dynamical core and physical processes that are consistent with BCC-AGCM2.1, but has a higher horizontal resolution. Results showed that both BCC-AGCM versions underestimated the global-mean total cloud cover (TCC), middle cloud cover (MCC) and low cloud cover (LCC), and that BCC_AGCM2.2 underestimated the global-mean high cloud cover (HCC). The global-mean cloud cover shows a systematic decrease from BCCA-GCM2.1 to BCC_AGCM2.2, especially for HCC. Geographically, HCC is significantly overestimated in the tropics, particularly by BCC_AGCM2,1, while LCC is generally overestimated over extra-tropical lands, but significantly underestimated over most of the oceans, especially for subtropical marine stratocumulus clouds. The leading EOF modes of CVS were extracted. The BCC_AGCMs perform well in reproducing EOF1, but with a larger variance explained. The two models also capture the basic features of EOF3, except an obvious deficiency in eigen- vector peaks. EOF2 has the largest simulation biases in both position and strength of eigenvector peaks. Furthermore, we investigated the effects of CVS on relative shortwave and longwave cloud radiative forcing (RSCRF and RLCRF). Both BCC_AGCM versions successfully reproduce the sign of regression coefficients, except for RLCRF in PC1. However, the RSCRF relative contributions from PC1 and PC2 are overestimated, while the relative contribution from PC3 is underes timated in both BCC_AGCM versions. The RLCRF relative contribution is underestimated for PC2 and overestimated for PC3.
基金Project was supported by the National Natural Science Foundation of China(Grant No.51541112).
文摘A comprehensive protective structure with rigidity and flexibility was put forward and designed in view of the quality and safety problems for the double vertical explosive welding of large titanium/steel cladding plate.The movement speed and displacement of the protective structure was calculated by establishing its physics model.The dynamics and stabilization properties were analyzed,and the protective structure parameters were optimized and devised.The comprehensive protective structure,which is composed of rigidity unit and flexibility wall,can bear the impact of detonation wave and the high-speed movement of the cladding plate.There are no damage and deformation in the protective structure and the cladding plate.The protective structure can be used many times.The bonding rate of the Ti/steel plate obtained was nearly 100%,and there is no deformation,surface cracks,and big wave and micro-defects.Therefore,the protective problems of the double vertical explosive welding can be solved effectively by the protective structure.
基金by the National Natural Science Foundation of China under grant No. 50775050the State Key Laboratory of Solidif ication Processing in NWPU (200702)
文摘The solidified structure of the thin-walled and complicated Ti-6AI-4V castings produced by the vertical centrifugal casting process was studied in the present work. The results show that the wall thickness of the section is featured with homogeneously distributed fine equiaxial grains, compared with the microstructure of the thick-walled section. The grain size of the castings has a tendency to decrease gradually with the increasing of the centrifugal radius. The inter-lamellar space in thick-walled casting parts is bigger than that of the thin-walled parts, and the profile of inter-lamellar space is not susceptible to the centrifugal radius.
基金This work is supported financially by the National Key R&D Program of China[2018YFC1406000,2016YFE0202700]Supported by the National Natural Science Foundation of China(NSFC)[Grant Nos.51809061,51639004]+1 种基金Supported by the Natural Science Foundation of Heilongjiang Province of China[LC2018021]Supported by the Fundamental Research Funds for the Central Universities[HEUCFM180111].
文摘In the ice-covered oceanic region,the collision between sea ice and offshore structures will occur,causing the crushing failure of ice and the vibration of structures.The vibration can result in fatigue damage of structure and even endanger the crews’health.It is no doubt that this ice-structure interaction has been noted with great interest by the academic community for a long time and numerous studies have been done through theoretical analysis,experimental statistics and numerical simulation.In this paper,the bond-based Peridynamics method is applied to simulate the interaction between sea ice and wide vertical structures,where sea ice is modeled as elastic-plastic material,with a certain yield condition and failure criterion.Oscillation equation of single-degree-of-freedom is considered to investigate the vibration features of the structure during the interaction process.The damage of ice,ice forces and vibration responses of structure in the duration are obtained through numerical simulation.A parametric investigation is undertaken to identify the key parameters,such as ice thickness,the diameter of structure and relative velocity that trigger the ice crushing,ice forces and vibration responses of the structure.Results indicate that all three parameters have a positive correlation with the overall level of ice force and vibration displacement.Besides,a velocity coefficient is proposed to predict the vibration displacement based on its relation with ice speed.
基金supported by the National Natural Science Foundation of China (Grant No. 41721004)
文摘This study investigates the vertical structure of variability in the cross-equatorial flows(CEFs)over the Maritime Continent and Indian Ocean in boreal summer,based on three reanalysis datasets:ERA-Interim,JRA-55 and NCEP-2.The results show a high consistency in the interannual variability among the reanalysis datasets,especially between ERAInterim and JRA-55,while great uncertainty exists in the decadal or long-term changes of CEFs.Further analyses on the interannual variability in CEFs indicate that there is a significant negative relationship between the upper-and lower-level CEFs over the Maritime Continent—that is,the northerlies at the upper level and southerlies at the lower level are both enhanced or weakened.This seesaw pattern is also significantly related to the CEFs over the Indian Ocean at the upper level and lower level(i.e.,the Somali jet).This close relationship between the upper-and lower-level CEFs and between the Maritime Continent and Indian Ocean is manifested as the leading mode of equatorial meridional winds in the vertical-zonal section over the Maritime Continent and Indian Ocean.Finally,it is found that ENSO is closely related to the vertical leading mode,and to all the CEFs at the upper and lower levels over the Maritime Continent and Indian Ocean.
基金supported by the National Natural Science Foundation of China(Grant Nos.51079082 and 51679132)the Nature Science Foundation of Shanghai City(Grant No.14ZR1419600)the Research Innovation Projects of 2013 Shanghai Postgraduate(Grant No.20131129)
文摘With the method of separation of variables and the eigenfunction expansion employed, an analytical solution is presented for the radiation and diffraction of a rectangular structure with an opening near a vertical wall in oblique seas, in which the unknown coefficients are determined by the boundary conditions and matching requirement on the interface. The effects of the width of the opening and the angle of incidence on the hydrodynamic characteristics of a rectangular structure with an opening near a vertical wall are mainly studied. The comparisons of the calculation results with wall-present and with wall-absent are also made. The results indicate that the variation trends of the heave added mass and excitation force with wall-present are almost the same as those with wall-absent, and that the peak values in the former case are obviously larger than those in the latter due to the reflection of the vertical wall.
基金Project supported by the National Natural Science Foundation of China (No.10176011).
文摘A new type of vibration structure (i.e. supporting system, called swing frame cus- tomarily) of vertical dynamic balancing machine has been designed, which is based on an analysis for the swing frame of a traditional double-plane vertical dynamic balancing machine. The static unbalance and couple unbalance can be e?ectively separated by using the new dynamic balancing machine with the new swing frame. By building the dynamics model, the advantages of the new structure are discussed in detail. The modal and harmonic response are analyzed by using the ANSYS7.0. By comparing the ?nite element modal analysis with the experimental modal analy- sis, the natural frequencies and vibration modes are found. There are many spring boards in the new swing frame. Their sti?nesses are di?erent and assorted with each other. Furthermore, there are three sensors on the measuring points. Therefore, the new dynamic balancing machine can measure static unbalance and coupling unbalance directly, and the interaction between them is faint. The result shows that the new vertical dynamic balancing machine is suitable for inertial measurement of ?ying objects, and can overcome the shortcomings of traditional double-plane vertical dynamic balancing machines, which the e?ect of plane-separation is inferior. The vertical dynamic balancing machine with the new vibration structure can ?nd wide application in the future. The modelling and analysis of the new vibration structure will provide theoretical basis and practical experience for designing new-type vertical dynamic balancing machines.
基金Supported by the Knowledge Innovation Program of Chinese Academy of Sciences (No.KZCX-3W-222 KZCX2-YW-Q11-02)+1 种基金National Basic Research Program of China (No.2007CB411802 2006CB403601)
文摘Empirical Orthogonal Function (EOF) analysis is used in this study to generate main eigenvector fields of historical temperature for the China Seas (here referring to Chinese marine territories) and adjacent waters from 1930 to 2002 (510 143 profiles). A good temperature profile is reconstructed based on several subsurface in situ temperature observations and the thermocline was estimated using the model. The results show that: 1) For the study area, the former four principal components can explain 95% of the overall variance, and the vertical distribution of temperature is most stable using the in situ temperature observations near the surface. 2) The model verifications based on the observed CTD data from the East China Sea (ECS), South China Sea (SCS) and the areas around Taiwan Island show that the reconstructed profiles have high correlation with the observed ones with the confidence level 〉95%, especially to describe the characteristics of the thermocline well. The average errors between the reconstructed and observed profiles in these three areas are 0.69℃, 0.52℃ and 1.18℃ respectively. It also shows the model RMS error is less than or close to the climatological error. The statistical model can be used to well estimate the temperature profile vertical structure. 3) Comparing the thermocline characteristics between the reconstructed and observed profiles, the results in the ECS show that the average absolute errors are 1.5m, 1.4 m and 0.17~C/m, and the average relative errors are 24.7%, 8.9% and 22.6% for the upper, lower thermocline boundaries and the gradient, respectively. Although the relative errors are obvious, the absolute error is small. In the SCS, the average absolute errors are 4.1 m, 27.7 m and 0.007℃/m, and the average relative errors are 16.1%, 16.8% and 9.5% for the upper, lower thermocline boundaries and the gradient, respectively. The average relative errors are all 〈20%. Although the average absolute error of the lower thermocline boundary is considerable, but contrast to the spatial scale of average depth of the lower thermocline boundary (165 m), the average relative error is small (16.8%). Therefore the model can be used to well estimate the thermocline.
基金The National Natural Science Foundation of China under contract Nos 41705081 and 41906005the Innovation Special Zone Project under contract No.18-H863-05-ZT-001-012-06the Open Project Fund of the Laboratory for Regional Oceanography and Numerical Modeling,Pilot National Laboratory for Marine Science and Technology(Qingdao)under contract No.2019A05.
文摘The sensitive area of targeted observations for short-term(7 d)prediction of vertical thermal structure(VTS)in summer in the Yellow Sea was investigated.We applied the Conditional Nonlinear Optimal Perturbation(CNOP)method and an adjoint-free algorithm with the Regional Ocean Modeling System(ROMS).We used vertical integration of CNOP-type temperature errors to locate the sensitive areas,where reduction of initial errors is expected to yield the greatest improvement in VTS prediction for the selected verification area.The identified sensitive areas were northeast−southwest orientated northeast to the verification area,which were possibly related to the southwestward background currents.Then,we performed a series of sensitivity experiments to evaluate the effectiveness of the identified sensitive areas.Results show that initial errors in the identified sensitive areas had the greatest negative effect on VTS prediction in the verification area compared to errors in other areas(e.g.,the verification area and areas to its east and northeast).Moreover,removal of initial errors through deploying simulated observations in the identified sensitive areas led to more refined prediction than correction of initial conditions in the verification area itself.Our results suggest that implementation of targeted observation in the CNOP-based sensitive areas is an effective method to improve short-term prediction of VTS in summer in the Yellow Sea.
文摘A new type of vibration structure of vertical dynamic balancing machine isdesigned, which is based on the analysis for swing frame of a traditional vertical dynamic balancingmachine. The static unbalance and couple unbalance can be separated effectively by using the newmachine with the new swing frame. By building the dynamics model, the advantages of the newstructure are discussed in detail. The modal and harmonic response are analyzed by using theANSYS7.0. By comparing the finite element modal analysis with the experimental modal analysis, thenatural frequencies and vibration modes are found out. There are many spring boards in the new swingframe. Their stiffness is different and assort with each other. Furthermore, there are threesensors on the measurement points. Therefore, the new dynamic balancing machine can measure thestatic unbalance and couple unbalance directly, and the influence between them is faint. The newstructure has the function of belt-strain compensation to improve the measurement precision. Thepractical result indicates that the new vertical dynamic balancing machine is suitable for inertialmeasurement of flying objects, and can overcome the shortcomings of traditional double-planevertical dynamic balancing machines. The vertical dynamic balancing machine with the new vibrationstructure can be widely used in the future applications. The modeling and analysis of the newvibration structure provide theoretic instruction and practical experience for designing new type ofvertical dynamic balancing machines. Based on the design principles such as stiffness-matching,frequency-adjacence and strain-compensation and so on, various new type of vibration structures canbe designed.
文摘In the coastal ecosystems of the Black Sea,macrophytobenthos and,in particular,the association of Cystoseira crinite,C.barbara,Cladostephus verticillatus,and Corallina mediterranea,with its thick vegetative canopy(VC),is the key contributor to primary production(PP).Though the vertical structure of the canopy,formed by the algal association,is of principal importance to the PP level,this subject has been long-neglected by researchers.The goal of our work was to compare vertical structures of the vegetative canopy of Cystoseira brown algae under diverse hydrodynamical conditions of the Crimean Peninsula coast.Samples were collected using the 50 cm×50 cm counting frame at eight stations positioned in shallow(55-60 cm deep)sites of Sevastopol Bay(Crimean Peninsula).Dry weight biomass of the VC was determined for all algae assemblage and for each algal species individually,per horizontal surface unit,at each height(Z).The study shows that:1)the VC is characterized by unimodal vertical distribution of biomass,with maximum estimate in the lower part,where the biomass increases to 85%of the total biomass;2)a series of single-peaked curves reliably describes the unimodal distribution of the biomass;thalli of different age groups are found along the canopy profile;and 3)algae found in epiphytic synusia prefer inhabiting the upper part of the VC.The role of environmental factors(seawater turbulence and solar radiation)is discussed in reference to the formation of the vertical structure,made up of the associations of the brown algae Cystoseira.
文摘The algebraic solitary wave and its associated eigenvalue problem in a deep stratified fluid with a free surface and a shallow upper layer were studied. And its vertical structure was examined. An exact solution for the derived 2D Benjamin-Ono equation was obtained, and physical explanation was given with the corresponding dispersion relation. As a special case, the vertical structure of the weakly nonlinear internal wave for the Holmboe density distribution was numerically investigated, and the propagating mechanism of the internal wave was studied by using the ray theory.
基金National Natural Science Foundation of China(41076118)Young Scientists Fund of National Natural Science Foundation of China Grant(41005018)
文摘Cloud profiling radar(CPR) onboard Cloud Sat allows for deep penetration into dense clouds/precipitation. In this study, tropical cyclones(TCs) are classified into three stages as developing, mature, and decaying. The circular TC area with the radius of 500 km is divided into five regions. The vertical structure characteristics of 94 Western Pacific TCs at different stages in different regions from June 2006 to February 2014 are statistically quantified using the Cloud Sat tropical cyclone overpass product(the CSTC Product). Contoured frequency by altitude diagrams(CFADs) of radar reflectivity show an arc-like feature and exhibit opposite distributions with a boundary at 5 km. Bright bands are found at this altitude, indicating melting layers. Deep convective(DC) clouds have the largest occurrence probability in the inner region, while Ci clouds occur more frequently in the outer region at 10-15 km. As clouds have the second largest vertical scale after DC clouds. Distributions of Ac, Cu, and Ns clouds at different stages have few distinctions.As the altitude increases, the ice effective radius and the distribution width parameter decrease while the particle number concentration increases. Moist static energy(MSE), cloud thickness(CT), liquid water path(LWP), ice water path(IWP), water vapor(WV), and rain rate(RR) all diminish along the radial direction and are significantly larger at the mature stage. The average value of MSE at the developing stage is larger than that at the decaying stage.
文摘In this paper, on the basis of the heat conduction equation without consideration of the advection and turbulence effects, one-dimensional model for describing surface sea temperature ( T1), bottom sea temperature ( Tt ) and the thickness of the upper homogeneous layer ( h ) is developed in terms of the dimensionless temperature θT and depth η and self-simulation function θT - f(η) of vertical temperature profile by means of historical temperature data.The results of trial prediction with our one-dimensional model on T, Th, h , the thickness and gradient of thermocline are satisfactory to some extent.
基金The China-Korea cooperative project"The study on the impact of the Yellow Sea Cold Water Mass to the ecosystem"under contract No.PE99165promoted by the Korea Institute of Ocean Science and Technology
文摘Using the hydroacoustic method with a 200 kHz scientific echo sounding system, the diel vertical migration (DVM) of the sound-scatteringlayer (SSL) in the Yellow Sea Bottom Cold Water (YSBCW) of the southeastern Yellow Sea was studied in April (spring) and August (summer) of 2010 and 2011. For each survey, 13-27 hours of acoustic data were continuously collected at a stationary station. The acoustic volume scattering strength (Sv) data were analyzed with temperature profile data. In the spring of both 2010 and 2011, the SSL clearly showed the vertical migration throughout the entire water column, moving from the surface layer at night to near the bottom during the day. Conductivity, temperature, and depth data indicated that the entire water column was well mixed with low temperature of about 8℃. However, the SSL showed different patterns in the summers of 2010 and 2011. In the summer of 2010 (≈28℃ at the surface), the SSL migrated to near the bottom during the day, but there were two SSLs above and below the thermocline at depth of 10-30 m at night. In the summer of 2011 (≈20℃ at the surface), the SSL extended throughout the entire water column at night, possibly owing to an abrupt change in sea weather conditions caused by the passage of a Typhoon Muifa over the study area. It was cancluded that the DVM patterns in summer in the YSBCW area may be greatly influenced by a strengthened or weakened thermocline.
基金supported by the National Natural Science Foundation of China (Grant No. 42105093 and 41975018)the China Postdoctoral Science Foundation (Grant No. 2020M670420)the Special Research Assistant Project。
文摘Turbulence in the nocturnal boundary layer(NBL)is still not well characterized,especially over complex underlying surfaces.Herein,gradient tower data and eddy covariance data collected by the Beijing 325-m tower were used to better understand the differentiating characteristics of turbulence regimes and vertical turbulence structure of urban the NBL.As for heights above the urban canopy layer(UCL),the relationship between turbulence velocity scale(VTKE)and wind speed(V)was consistent with the“HOckey-Stick”(HOST)theory proposed for a relatively flat area.Four regimes have been identified according to urban nocturnal stable boundary layer.Regime 1 occurs where local shear plays a leading role for weak turbulence under the constraint that the wind speed V<VT(threshold wind speed).Regime 2 is determined by the existence of strong turbulence that occurs when V>VT and is mainly driven by bulk shear.Regime 3 is identified by the existence of moderate turbulence when upside-down turbulence sporadic bursts occur in the presence of otherwise weak turbulence.Regime 4 is identified as buoyancy turbulence,when V>VT,and the turbulence regime is affected by a combination of local wind shear,bulk shear and buoyancy turbulence.The turbulence activities demonstrated a weak thermal stratification dependency in regime 1,for which within the UCL,the turbulence intensity was strongly affected by local wind shear when V<VT.This study further showed typical examples of different stable boundary layers and the variations between turbulence regimes by analyzing the evolution of wind vectors.Partly because of the influence of large-scale motions,the power spectral density of vertical velocity for upsidedown structure showed an increase at low frequencies.The upside-down structures were also characterized by the highest frequency of the stable stratifications in the higher layer.
文摘We investigated the vertical variability of toxic potential (i.e. proportions of cells containing microcystin genes) and genotypic structure within different populations of Microcystis aeruginosa that developed in deep artificial reservoirs on the Loire River (France). We demonstrated that a great qualitative vertical heterogeneity could exist within a single bloom of this cyanobacterium in deep lakes. Indeed, we observed important vertical shifts of both toxic potential and genotypic structure, whatever the bloom magnitude. These variations occurred mainly within the euphotic zone and proved to occur independently from abundance vertical shifts. One of the most striking results of this study is that the genotypic structure of a population of M. aeruginosa was more variable between different depths sampled at a single site than between different sites of the same reservoir sampled on top of the water column. In the same way the proportion of potentially toxic cells was sometimes more variable vertically than horizontally. The occurrence of such vertical heterogeneity in three different blooms suggests that this could be a frequent pattern within populations of M. aeruginosa.