Effects of drought on non-structural carbohydrates and C,N,and P stoichiometric characteristics of Pinus yunnanensis seedlings

To study non-structural carbohydrate character-istics and nutrient utilization strategies of Pinus yunnanen-sis under continuous drought conditions,2-year-old seed-lings were planted in pots with appropriate water,light and moderate and severe drought treatments[(80±5),(65±5),(50±5),and(35±5)%of field water-holding capacity].Non-structural carbohydrates,carbon(C),nitrogen(N),and phosphorus(P)concentrations were measured in each plant component.The results show that:(1)With increasing drought,non-structural carbohydrates gradually increased in leaves,stems,and coarse roots,while gradually decreased in fine roots;(2)C concentrations of all were relatively stable under different stress levels.Phosphorous utilization of each component increased under light and moderate drought conditions,while N and P utilization efficiency of each plant component decreased under severe drought.Growth was mainly restricted by N,first decreasing and then increasing with increased drought;(3)There was a correlation between the levels of non-structural carbohydrates and C,N,and P in each component.Changes in N concentration affected the interconversion between soluble sugar and starch,which play a regulatory role in the fluctuation of the concentration of non-structural carbohydrates;and,(4)Plasticity analysis showed that P.yunnanensis seedlings responded to drought mainly by altering starch concentration,the ratio of soluble sugar to starch in leaves and stems,and further by alter-ing N and P utilization efficiencies.Overall,these results suggest that the physiological activities of all organs of P.yunnanensis seedlings are restricted under drought and that trade-offs exist between different physiological indicators and organs.Our findings are helpful in understanding non-structural carbohydrate and nutrient adaptation mechanisms under drought in P.yunnanensis seedlings.

Molecular Dissection of Relationship Between Seedling Characteristics and Seed Size in Rice

Seedling characteristics such as leaf emergency, chlorophyll content, and height are important for early growth and also associated with seed size. Quantitative trait loci (QTLs) for rice (Oryza sativa L.) seedling characteristics (leaf length, chlorophyll content, seedling height) and seed size were identified with an attempt to explore the relationship between seedling characteristics and seed size, using a recombinant inbred population derived from a cross between Zhenshan 97A and Minghui 63. Two, one, five, four, four, and nine QTLs were detected for chlorophyll a content, total chlorophyll, length of the second seedling leaf, length of the third seedling leaf, seedling height, and seed size, respectively. The results indicated that four QTLs for seed size and four QTLs for seedling characteristics shared several similar regions (G359-RG532, C567-RG236, RZ403-R19, C371-C405a), respectively, suggesting a close association between seedling characteristics and seed size. Several chromosomal regions influencing seed size, however, had no effects on seedling characteristics, suggesting that it was possible to improve seedling vigor without increasing grain size.

Effects of soil moisture and light intensity on ecophysiological characteristics of Amorpha fruticosa seedlings

We investigated the combined effects of soil moisture and light intensity on the growth, development and ecophysiological characteristics of one-year old Amorpha fruticosa seedlings. Soil moisture and light intensity influenced the ecophysiological characteristics of Amorpha fruticosa seedlings. Soil moisture resulted in the decreases of growth rate, individual size, net photosynthetic rate, transpiration rate, leaf water loss rate （WLR）, and biomass accumulation of plant parts, and led to increased leaf water saturation deficit （WSD）. Under water stress, more photosynthetic products were allocated to root growth. With decreasing light intensity, net photosynthetic rate, transpiration rate, chla/b, water saturation deficit, water use efficiency, water loss rate and biomass accumulation declined, while Chla, Chlb, Chla＋b and carotenoids （Car） increased and more photosynthetic products were allocated to stem and leaf growth. Maximum growth vigor, net photosynthetic rate and total biomass accumulation in Amorpha fruticosa seedlings was recorded at 75 80% soil water-holding capacity and 100% light density in greenhouse environments.

Effects of sodium benzoate on growth and physiological characteristics of wheat seedlings under compound heavy metal stress

In this study, we investigated the effect of exogenous sodium benzoate on wheat seedlings(Yangmai 16) grown under heavy metal stress. The results showed that 2.4 mmol kg-1 of heavy metals significantly inhibited growth and delayed emergence of wheat seedlings. Under compound heavy metal stress, application of 2-4 g L^-1 sodium benzoate significantly increased(P<0.01) chlorophyll content and chlorophyll fluorescence parameters Fv/Fm and Fv/Fo of wheat, compared to the control(water treatment). Further analysis showed that application of 2-4 g L^-1 sodium benzoate alleviated osmotic stress by promoting the accumulation of osmolytes such as soluble proteins and free proline, increased the activity of superoxide dismutase(SOD) and reduced malondialdehyde content(MDA). In contrast, higher concentrations of sodium benzoate solution(>6 g L^-1) inhibited the growth of wheat seedlings and even caused damage to seedlings. Correlation analysis showed that when the sodium benzoate concentration was in the range of 1.97-3.12 g L^-1(2016) and 1.58-3.27 g L^-1(2017), values of chlorophyll and its components, root activity, SOD activity, soluble protein, and free proline content were the highest. When the sodium benzoate concentration was raised to 2.59 g L^-1(2016) or 3.02 g L^-1(2017), MDA content was the lowest. Ultimately, exogenous sodium benzoate(2-4 g L^-1) facilitates root development and improves the root activity of wheat seedlings grown under compound heavy metals stress, thereby effectively alleviating the damage of compound heavy metal stress in wheat seedlings.

Effect of La-Gly on Physiological and Biochemical Characteristics of Glycine max Seedling under Pb Stress

The physiological and biochemical characteristics of Glycine max seedlings were changed under the high concentration of Pb stress, in which the malony dialdehyde (MDA) content, plasmalemma, activities of catalase (CAT) and peroxidase (POD) were increased drastically. The damage effects of Pb were reduced when 15 mg . L-1 La-Gly was used to spray Glycine max seedlings. The results of La-Gly treatment show that the effects are related to the decrease of Pb content in plants, the increase of photosynthetic rate and nitrate reductase (NR) activity, decrease of MDA content and cell membrane permeability, and maintenance of activities of CAT and POD.

Growth Performance of Phoebe bournei Generations in Seedling Stage

[Objective] The aim was to explore growth performance of Phoebe bournei in seedling stage.[Method] Based on 17 superior plus trees of Phoebe bournei,a seedling cultivation test was conducted in Jindong Forest Farm in Yongzhou,Hunan Province,exploring growth indices and analyzing seedling height and ground diameter.[Result] As per the selection rate at 20％,PB17,PB9 and PB12 were selected from the 17 species.Specifically,the seedling height averaged 36.52 cm of the selected three seedlings,which was higher by 16.1％ compared with average of 17 species （31.45％）.As for ground diameter,the selected species averaged 2.6 mm,which was higher （13.0％） compared with the average level （2.3 mm）.[Conclusion]Both of seedling height and ground diameter reached extremely significant differences,which indicated that half-sib families of Phoebe bournei have rich variations,with selection and breeding potential.

Review on the proceeding of automatic seedlings classification by computer vision

The classification of seedlings is important to ensure the viability of seedlings after transplantation and is acknowledged as a key factor in forestation and environmental improvement. Based on numerous papers on automatic seedling classification (ASC), the seedling grading theory, traditional grading methods, the background and the proceeding of ASC techniques are described. The automation of the measurement of seedling morphological characteristics by photoelectric meters and computer vision is studied, and the automatic methods of the current grading systems are described respectively. And the further researches on ASC by computer vision are proposed.

Effects of light intensity on leaf microstructure and growth of rape seedlings cultivated under a combination of red and blue LEDs

The aim of this study was to evaluate the growth of rape （Brassica napus L.） seedlings under different light intensities to select appropriate conditions for cultivation in an indoor system. Seedlings were grown under different light intensities of red and blue light provided by light-emitting diodes （LEDs） and their self-adjustment ability and changes in leaf microstructure were evaluated. Light was supplied by red LEDs with peak wavelengths of 630 （R1） and 660 nm （R2） and by blue LEDs （B） with a peak wavelength of 445 nm （the light intensity ratio of R1：R2：B was 3：3：2）, at intensities of 400 （R1R2B400）, 300 （R1R2B300）, and 200 μmol m-2 s-1 （R1R2B200）. Natural solar light served as the control （C）. Plant height, stem diameter, root length, leaf area, and dry weight of rape seedlings gradually increased with increasing light intensity. The seedlings in the R1R2B400 treatment grew more vigorously, while those in the R1R2B200 treatment were weaker. The photosynthetic pigment contents did not differ significantly between the R1R2B400 treatment and C, but were significantly lower in the R1R2B300 and R1R2B200 treatments. The highest intercellular CO2concentration, stomatal conductance, and transpiration rate were in the R1R2B300 treatment. The highest photosynthetic rate was in the R1R2B400 treatment, and was related to more compact leaves, thicker and tidier palisade and spongy tissues, and well-developed chloroplasts. In contrast, the seedlings in the R1R2B200 treatment had disordered mesophyll cells, round chloroplasts, and fractured and fuzzy grana lamellae, all of which inhibited plant growth. In conclusion, the seedlings in the R1R2B400 treatment had well-developed leaves, which favored photosynthesis. Compared with the light intensities below 300 μmol m-2 s-1, the light intensity of 400 μmol m-2 s-1 provided by a cembination of red and blue LEDs was beneficial for cultivating strong and healthy rape seedlings in an artificial system.

The response of Caragana microphylla seedlings to water table changes in Horqin Sandy Land,China

This paper focuses on the growth response of Caragana microphylla seedlings to changes of artificially controlled water table in Horqin Sandy Land, China. Monitoring results of soil water content shows that soil moisture is closely correlated to groundwater depths. Soil volumetric water increased rapidly when close to water sources and finally stabilized in a saturated state. The soil moisture trend of CK （control） increased gradually at 0-50 cm of soil depth then decreased to 4% below 50 cm soil depth. C. microphylla can adapt to different soil environments by changes in ecological and physiological characteristics. By comparing the ecological characteristics of C. microphylla seedlings at various water tables, we found that a shallow water table of 40 cm depth inhibited seedling groundwater depth of 120 em was more advantageous for plant growth because of weak ecological characteristics. A height and canopy growth of C. microphylla seedlings. During the first two years, the most suitable water depth for root biomass was 120 cm, and 180 cm for root length. The growth of vertical roots is positively correlated with groundwater depth, and root thickness is the determinate factor for root biomass while the fine root is the determinate factor for root length. A thick root would grow much more in a natural drought environment while access to ground water promotes the growth of fine roots.

Influence of topography,soil properties and plant community on the biomass of Abies georgei var.smithii seedlings in Southeast Tibet

Biomass of seedlings at different developing stages of growth is important information for studying the response of species to site conditions.The objectives of this study was to explore the distribution characteristics of AGB(above-ground biomass)and BGB(below-ground biomass)of Abies georgei var.smithii seedlings of different ages,and investigate the effects of topography(slope aspect,altitude),plant community characteristics(crown density,species diversity,etc.),and soil properties(soil physical and chemical properties)on the biomass and its allocation.Seedlings in five age classes(1–2,3–4,5–6,7–8,and 9–10 years old)were collected by full excavation from 6 elevations(3800 m,3900 m,4000 m,4100 m,4200 m,4300 m)on the north and south slopes of Sejila Mountain in Tibet.15seedlings of each age class were investigated at one altitude.The individual effects of seedling age(SA)and the interaction effects of SA,slope aspect(SL),and elevation(EG),namely,SL×EG,SL×SA,EG×SA,and SL×EG×SA,had significant effects on the AGB of the seedlings(p<0.05),whereas BGB was only significantly affected by SA(p<0.001).The AGB and BGB of the seedlings showed a binomial growth trend with the increase in seedling age,and had an allometric relationship at different elevations,α(allometric exponential)varied from 0.913 to 1.046 in the northern slope,and from 1.004 to 1.268 in the southern slope.The biomass of seedlings on the northern slope was remarkably affected by stand factors,with a contribution rate of 47.8%,whereas that on the southern slope was considerably affected by soil factors with a contribution rate of 53.2%.The results showed that age was the most important factor affecting seedling biomass.The allometric pattern of seedling biomass was relatively stable,but in a highaltitude habitat,A.georgei var.smithii seedlings increased the input of BGB.Understanding seedling biomass allocation and its influencing factors is useful for evaluating plants’ability to acquire resources and survival strategies for adaptation to the environment in Tibet Plateau.

TIB for Micropropagation and the Relationship between Anthocyanins and Chlorophyll of Strawberry Seedlings

In this study, we employed a temporary immersion bioreactor(TIB)system for the micropropagation of strawberry seedlings. The TIB method and the conventional method for the micropropagation of strawberry seedlings were compared in terms of explant propagation coefficient, seedling fresh weight,contents of anthocyanins and chlorophyll, and photosynthetic characteristics. The results showed that an inoculation density of 40 explants/L was suitable for the micropropagation of ’Benihope’ strawberry seedlings in the TIB. The propagation coefficient, fresh weight, photosynthetic rate, stomatal conductance, transpiration coefficient, and the contents of total anthocyanins and chlorophyll of seedlings micropropagated in the TIB were significantly higher than those of the seedlings micropropagated by the conventional method. In conclusion, the TIB method was superior to the conventional method in the micropropagation of strawberry seedlings.The negative reciprocity did not occur between anthocyanins and chlorophyll in strawberry seedlings cultured under forced ventilation such as TIB.

Evaluation and Selection of Date Palm Varieties from Seedling Progeny in Saudi Arabia

A survey was conducted in 2011 in the major date growing regions of Saudi Arabia to identify and evaluate date palms with fruit characteristics preferred by the producers and consumers. The regions in the Kingdom surveyed for this purpose were Al-Zulfi, Al-Hassa, Najran, Al-Madinah Al-Munawarah and AI-Qassim. Seventy palms from seedlings were initially identified for the study based on the following fruit characteristics, i.e., fruit length, breadth, weight, color, taste, texture, sugar content, external appearance and also the market value. Subsequently, 12 palms （Al-Afiah, Duhiba, Adbah, Aliah, Shamshula, Masifat Al-Asedy, Suwaid, Batlyah, AI-Nassar, Hussan, Sukariyat Al-Zulfi and Al-Hamat） mainly from the Zulfi and Riyadh regions were found to meet the required standards with potential for commercial date farming. Among these, fruits of Adbah had the highest total sugars of 72.9% followed by Duhiba with a total sugar content of 71.9%. Fruit length was maximum （49.1 mm） in the variety Allah, while the fruits of the variety Hussan recorded the maximum weight of 19.5 g. Furthermore, it is pertinent to mention that fruit size of the 12 selected cultivars was generally large and comparable to Majdool, a well-known date palm cultivar of North Africa. The fruits of the selected cultivars were also tasty and can be stored separately.

Response of Somatal Characteristics and its Plasticity to Different Light Intensities in Leaves of Seven Tropical Woody Seedlings

Stomatal characteristics and its plasticity were surveyed in leaves of four canopy species, Shoreachinensis, Pometia tomentosa, Anthocephalus chinensis, Calophyllun polyanthum and three middle-layerspecies, Barringtonia pendula, Garcinia hanburyi, Horsfieldia tetratepala, acclimated to different lightconditions for more than one year. All plants stomata are distributed on the abaxial of leaves. Pometiatomentosa and Barringtonia pendula have higher stomatal density and the guard cell length of Anthocephaluschinensis and Calophyllun polyanthum were much greater than others. Stomatal density and stomatal index(ratio of stomatal numbers to epidermal cell number) were increased with growth irradiance increased, whilenumbers of stomata per leaf were higher in the low than the high relative PFD, and stomatal conductance ofleaves was the highest in the 50% of full light except for Anthocephalus chinensis. The relative PFD has littleeffects on the guard cell length of all seven plants. We have also found a significant negative correlationbetween stomatal density and leaf area, but the stomatal conductance was not significantly positive with thestomatal conductance. The analysis of phenotypic plasticity of stomatal characteristics showed: plasticityindex for stomatal index and numbers of stomatal per leaf were similar for canopy and middle-layer species,while the plasticity index of stomatal density and stomatal conductance were significantly greater for canopyspecies than middle-layer species. The high plasticity of canopy species was consistent with the hypothesisthat specialization in a more favorable environment increases plasticity.

Phytoremediation Characteristics of Pb and Cd Contaminated Soils by Ginkgo Seedling

In this article,we investigated effects of lead and cadmium on the biomass of ginkgo seedlings, and roles of the seedlings in accumulation characteristics of the heavy metals and remediation of the soil contaminated by Cd2+ or Pb2+ as well as their combination.The results showed that both Cd2+ and Pb2+ significantly or very significantly reduced biomass increment of ginkgo seedlings,with Pb2+ more adverse than Cd2+.The Cd2+-Pb2+ compound pollution further reduced biomass increment of ginkgo seedlings.The mass fractions and enrichment coefficient of Pb2+ and Cd2+ in ginkgo seedlings were in an order of root】stem】leaf,which was in favor of transportation Pb2+ and Cd2+ from root to stem and leave.The Cd2+-Pb2+ compound pollution promoted ginkgo seedling absorption to Cd2+ and Pb2+.The capacity of the upward transport of Pb2+ was greater than that of Cd2+,which suggested an obvious dose-effect and synergistic effect.Cd2+ and Pb2+ accumulations in ginkgo seedlings were relatively large.The accumulation in roots was higher than that in stems and leaves.The ability of the absorption,transportation and accumulation of ginkgo seedlings to Pb2+ was relatively greater under Cd2+-Pb2+ compound pollution.The phytoremediation efficiency of Cd2+ and Pb2+ was relatively high.

NaCl胁迫下外源亚精胺对马铃薯幼苗生理特征的影响

【目的】研究NaCl胁迫下外源亚精胺(Spd)对马铃薯幼苗生理特征的影响,为新疆盐渍化区域马铃薯种植提供技术参考。【方法】以晋薯16号马铃薯幼苗为材料,采用不同浓度NaCl模拟盐胁迫处理,研究外源亚精胺Spd对马铃薯植株生长、叶片抗氧化酶活性、渗透调节物质含量的影响。【结果】盐胁迫抑制了马铃薯植株生长,使得超氧化物歧化酶(SOD)、过氧化物酶(POD)、过氧化氢酶(CAT)、抗坏血酸过氧化物酶(APX)等酶促类抗氧化酶活性表现为低盐胁迫促进,高盐胁迫抑制;抗坏血酸(ASA)、谷胱甘肽(DSH)等非酶促类抗氧化酶含量均降低;脯氨酸、可溶性糖、氨基酸含量升高;外源亚精胺(Spd)通过提高叶绿素含量、根系活力,调节渗透调节物质含量,增强抗氧化酶活性等方式维持植物体的正常生理代谢功能。【结论】外源亚精胺(Spd)可提升马铃薯耐盐性,缓解盐胁迫对马铃薯植株的抑制作用,其中0.9 mmol/L Spd(S 3)浓度缓解效果最明显。

不同基质对‘丹红紫叶’紫薇容器苗开花和光合特性的影响

[目的]为探究不同育苗基质对‘丹红紫叶’紫薇容器苗开花及光合特性的影响,筛选出紫薇容器苗培育的适宜育苗基质,为紫薇容器苗高效培育提供理论指导。[方法]以‘丹红紫叶’紫薇2年生容器苗为研究对象,用圃地土、泥炭、珍珠岩、蛭石不同比例配制16种育苗基质处理,以圃地土为对照(CK)基质,测定不同处理对容器苗开花及光合特性的影响。[结果]不同育苗基质对‘丹红紫叶’紫薇容器苗的开花及光合特性影响显著,T11处理的开花指标和光合特性指标综合表现最好,花期、花径、花序长和宽分别比CK增加了23d、22.63%、26.40%和30.20%,叶绿素相对含量、净光合速率、蒸腾速率、气孔导度、水分利用效率分别比CK显著提高了8.88%、413.58%、115.20%、100.00%、138.46%,T11处理的胞间CO 2浓度最低,比CK显著降低29.27%。开花指标与光合特性指标的相关性分析表明,花序与净光合速率、水分利用效率显著正相关,与胞间CO_(2)浓度显著负相关。主成分分析的综合得分最高为T11处理,最低为CK,T11处理基质配方有利于促进‘丹红紫叶’紫薇容器苗开花,提高光合效率。[结论]圃地土30%+泥炭50%+蛭石10%+珍珠岩10%的基质是‘丹红紫叶’紫薇2年生容器苗培育的最适宜基质。

不同夏花生品种苗期发育规律研究

以高油酸花生品种豫花37号(YH37)和普通油酸花生宛花2号(WH2)为研究材料,采用盆栽试验,调查不同品种苗期形态发育规律、叶片功能、激素含量等的变化,为制定精准科学的管理措施提供理论依据。结果表明,两个品种苗期发育规律不同,豫花37号苗期以地下部生长为主,至出苗后28 d,豫花37号地下部赤霉素、细胞分裂素和生长素含量分别为宛花2号的1.06、1.22和1.09倍;根长、根尖数、地下部干质量、根冠比分别较宛花2号高75.51%、95.03%、10.64%和29.84%。宛花2号苗期以地上部生长为主,至出苗后28 d,主茎高、侧枝长、叶片数、分枝数、地上部干质量分别较豫花37号高12.41%、65.00%、6.12%、7.69%和17.47%。与宛花2号相比,豫花37号叶片光转化能力由低到高逐渐升高,phi2值由97.47%升高到100.93%,且在整个生育期内豫花37号叶片的光保护能力更强,受光损伤的程度更小。基于此,在苗期豫花37号应适当补充中微量元素,以保证其地上地下部协调生长;宛花2号应减少叶面肥和促进茎叶生长的调理剂用量,从而达到壮苗的目的。

施用氯化钠对椰子幼苗生理特性的影响

为深入了解施用氯化钠(NaCl)对椰子幼苗生理特性的影响,通过施用不同量的NaCl,30 d后测定叶绿素含量、丙二醛(MDA)含量、可溶性糖含量、脯氨酸含量、超氧化物歧化酶(SOD)、过氧化物酶(POD)和过氧化氢酶(CAT)活性。结果显示:施用100 g和150 g NaCl时,叶绿素含量显著升高,MDA和可溶性糖含量明显降低,POD、SOD和CAT活性显著增加,脯氨酸含量少量增加;200 g NaCl处理条件下叶绿素含量显著降低,MDA和可溶性糖含量明显减少,SOD和CAT活性显著降低,脯氨酸含量显著增加。试验表明适量施用NaCl可以增强椰子幼苗光合作用,抑制膜脂过氧化,促进物质转化等。

光质对苜蓿芽苗菜生理特性和抗氧化酶活性的影响

研究了不同光质[红光、蓝光、红蓝混合光、白光(CK)]对苜蓿芽苗菜生理特性及抗氧化酶活性的影响。结果表明,红光可显著提高可溶性糖含量,蓝光和红蓝混合光则降低了可溶性糖含量;蓝光处理的可溶性蛋白质含量、根系活力、过氧化物酶(POD)活性显著高于其他光质处理;红蓝混合光处理的维生素C含量和叶绿素含量最高,且叶绿素含量与百株鲜重的拟合方程的决定系数(R2)为0.930 9;不同光质处理的超氧化物歧化酶(SOD)活性之间无显著差异。红蓝混合光处理的百株维生素C含量和百株可溶性蛋白质含量均较高,红蓝混合光是苜蓿芽苗菜生产中较好的补光光源。

胞外多糖对盐胁迫下荆芥种子萌发、幼苗生长及生理特性的影响

为提高盐渍地中荆芥的种植效益,探究植物乳杆菌(Lactobacillus plantarum)胞外多糖对盐胁迫下荆芥种子萌发、幼苗生长的影响,同时从幼苗根系生理特性上分析其具体作用机制,设置6个处理[空白对照组(CK1)、盐胁迫对照组(CK2)、50 mg/kg胞外多糖组(T1)、100 mg/kg胞外多糖组(T2)、200 mg/kg胞外多糖组(T3)、400 mg/kg胞外多糖组(T4)]进行试验。结果表明,胞外多糖对荆芥种子萌发和幼苗生长具有显著的促进作用。其中,添加200 mg/kg胞外多糖(T3)效果最显著,与CK2相比,其种子发芽率、发芽势、发芽指数和活力指数分别显著增加76.13%、144.16%、137.45%、265.28%,幼苗株高、茎粗、地下干质量和地上干质量分别显著升高60.49%、14.73%、40.08%、85.39%,脯氨酸、可溶性蛋白、可溶性糖含量分别提高21.02%、84.56%、40.02%,Na+含量降低54.63%,K+含量升高95.24%,Na+/K+值降低76.76%,根系电解质渗出率、丙二醛(MDA)含量分别降低44.54%、43.24%,超氧化物歧化酶(SOD)、过氧化氢酶(CAT)、过氧化物酶(POD)活性分别显著提高30.39%、34.59%、24.31%。综上所述,植物乳杆菌胞外多糖可以通过根系渗透调节和激活抗氧化酶防御系统来提高幼苗的抗盐性,进而促进荆芥种子萌发和幼苗生长。