Extreme climate has increasingly led to negative impacts on forest ecosystems globally,especially in semiarid areas where forest ecosystems are more vulnerable.However,it is poorly understood how tree growth is affect...Extreme climate has increasingly led to negative impacts on forest ecosystems globally,especially in semiarid areas where forest ecosystems are more vulnerable.However,it is poorly understood how tree growth is affected by different drought events.In 2006–2009,the larch plantations in the semiarid areas of Northwest China were negatively affected by four consecutive dry years,which was a very rare phenomenon that may occur frequently under future climate warming.In this study,we analyzed the effect of these consecutive dry years on tree growth based on the data of the tree rings in the dominant layer of the forest canopy on a larch plantation.We found that the tree-ring width index(RWI)in dry years was lower than that in normal years,and it experienced a rapidly decreasing trend from 2006 to 2009(slope=-0.139 year^(-1),r=-0.94)due to water supply deficits in those dry years.Drought induced legacy effects of tree growth reduction,and consecutive dry years corresponded with greater growth reductions and legacy effects.Growth reductions and legacy effects were significantly stronger in the third and fourth consecutive dry years than that of single dry year(p<0.05),which might have been due to the cumulative stress caused by consecutive dry years.Our results showed that larch trees experienced greater tree growth reduction due to consecutive dry years and their legacy effect,and the trees had lower recovery rates after consecutive dry years.Our results highlight that consecutive dry years pose a new threat to plantations under climate warming,and thus,the effect of climate extremes on tree growth should be considered in growth models in semiarid areas.展开更多
Larix gmelinii var.principis-rupprechtii(Mayr.)Pilger is an important native tree species in North China with advantages of fast growth,straight trunk,and good wood properties.The multi-year and multi-site breeding re...Larix gmelinii var.principis-rupprechtii(Mayr.)Pilger is an important native tree species in North China with advantages of fast growth,straight trunk,and good wood properties.The multi-year and multi-site breeding research of families of the species has not been reported previously.Based on diameter at breast height(DBH),height and volume of 25 families on four experimental sites,we calculated variance components,genetic parameters,juvenile and mature trait correlations and made genotype main effect plus genotype×environment interaction effect(GGE)biplot based on the breeding values estimated using the method of best linear unbiased prediction(BLUP).Compared with height,DBH and volume had higher heritability and larger variation coefficients,making them the more suitable traits for family selection and evaluation.Based on these,GGE biplots containing 20 combinations of site×age were drawn using data at 13 to 17 years when the interactions between family and location were strong.Test sites classifications based on DBH,and volume were inconsistent,with two categories for DBH and one for volume.The Guyuan site was the most suitable with strong discriminating ability,high representativeness and stability among tree ages.Integrating the ranking results of DBH and volume,families 66,76,82 and 111 were high-yielding and stable,families 78 and96 were high-yielding with above average stability,families72 and 79 were high-yielding with below average stability,whereas stability of family 100 was inconsistent between DBH and volume.Early selection based on DBH was convenient and reliable,and can be made at seven years.This study provides support for the selection of Larix gmelinii var.principis-rupprechtii families in Hebei province and an example for the application of stem analysis data from multiple sites in tree breeding.展开更多
Korean larch(Larix olgensis)is one of the main tree species for aff orestation and timber production in northeast China.However,its timber quality and growth ability are largely infl uenced by crown size,structure and...Korean larch(Larix olgensis)is one of the main tree species for aff orestation and timber production in northeast China.However,its timber quality and growth ability are largely infl uenced by crown size,structure and shape.The majority of crown models are static models based on tree size and stand characteristics from temporary sample plots,but crown dynamic models has seldom been constructed.Therefore,this study aimed to develop height to crown base(HCB)and crown length(CL)dynamic models using the branch mortality technique for a Korean larch plantation.The nonlinear mixed-eff ects model with random eff ects,variance functions and correlation structures,was used to build HCB and CL dynamic models.The data were obtained from 95 sample trees of 19 plots in Meng JiaGang forest farm in Northeast China.The results showed that HCB progressively increases as tree age,tree height growth(HT growth)and diameter at breast height growth(DBH growth).The CL was increased with tree age in 20 years ago,and subsequently stabilized.HT growth,DBH growth stand basal area(BAS)and crown competition factor(CCF)signifi cantly infl uenced HCB and CL.The HCB was positively correlated with BAS,HT growth and DBH growth,but negatively correlated with CCF.The CL was positively correlated with BAS and CCF,but negatively correlated with DBH growth.Model fi tting and validation confi rmed that the mixed-eff ects model considering the stand and tree level random eff ects was accurate and reliable for predicting the HCB and CL dynamics.However,the models involving adding variance functions and time series correlation structure could not completely remove heterogeneity and autocorrelation,and the fi tting precision of the models was reduced.Therefore,from the point of view of application,we should take care to avoid setting up over-complex models.The HCB and CL dynamic models in our study may also be incorporated into stand growth and yield model systems in China.展开更多
In order to understand the effects of thinning on microsite conditions and natural regeneration in the larch plantation, thinning experiment was conducted in a 40-year-old Larix olgensis plantation in Qingyuan County ...In order to understand the effects of thinning on microsite conditions and natural regeneration in the larch plantation, thinning experiment was conducted in a 40-year-old Larix olgensis plantation in Qingyuan County in eastern Liaoning Province, China in 2003-2004 Five thinning treatments (0%, 10.2%, 19.8%, 29.7% and 40.3% thinned) were designed on the same site. After thinning, canopy openness and the microsite conditions such as photosynthetic photon flux density (PPFD), soil moisture content, and soil temperature were measured in one growing season. Meanwhile, the investigation of natural regeneration was conducted at the end of the growing season. The results showed that the canopy openness increased with the increase of thinning intensities. PPFD and soil temperature and soil moisture content in different soil layers were positively relative with canopy openness after thinning. The richness of regenerating tree species did not significantly increase (p=0.30) after one growing season since thinning, but the regeneration density and frequency of tree species increased significantly (p〈0.05). In addition, the number of regenerating tree species increased, and the increment was correlated with the characteristics of iudividual tree species. The increasing percentage of regenerating seedlings of the shade-intolerant tree species was more than that of shade-tolerant tree species. Among the investigated regeneration species, the biggest response of seedling emergency to the canopy openness was Phellodendron amurense. This paper confirmed the following conclusions: after thinning, the variety of regenerating tree species was correlative with the characteristics of regenerating tree species, and the distribution of unthinned trees and the site conditions in the investigated larch plantation were the additional factors influencing, the regeneration.展开更多
Because growth ring data have temporal features, time series analysis can be used to simulate and reveal changes in the life of a tree and contribute to plantation management. In this study, the autoregressive(AR) and...Because growth ring data have temporal features, time series analysis can be used to simulate and reveal changes in the life of a tree and contribute to plantation management. In this study, the autoregressive(AR) and moving average modeling method was used to simulate the time series for growth ring density in a larch plantation with different initial planting densities. We adopted the Box–Jenkins method for the modeling, which was initially based on an intuitive analysis of sequence graphs followed by the augmented Dickey–Fuller stationarity test. The order p and q of the ARMA(p, q) model was determined based on the autocorrelation and partial correlation coefficient figure truncated on the respective order.Through the residual judgment, the model AR(2) was only fitted to the larch growth ring density series for the plantation with the 1.5 9 2.0 m^2 initial planting density.Because the residuals series for the other three series was not shown as a white noise sequence, the modeling was rerun. Larch wood from the initial planting density of2.0 9 2.0 m^2 was modeled by ARMA(2, 1), and ARMA((1, 5), 3) fitted to the 2.5 9 2.5 m^2 initial planting density,and the 3.0 9 3.0 m^2 was modeled by AR(1, 2, 5).Although the ARMA modeling can simulate the change in growth ring density, data for the different growth ring time series were described by different models. Thus, time series modeling can be suitable for growth ring data analysis, revealing the time domain and frequency domain of growth ring data.展开更多
Through 5 years of phenological observations on Larix principis-rupprechtii Mayr. in primary seed orchard and studies on population and individuals of clones, the annual periodic phenological laws were revealed and th...Through 5 years of phenological observations on Larix principis-rupprechtii Mayr. in primary seed orchard and studies on population and individuals of clones, the annual periodic phenological laws were revealed and the annual phe-nological periodic table was drawn up. The correlation between various phenophases, the air temperature and active accumu-lated temperature were analyzed and expounded. The authors also analyzed the similarities and differences of phenophases among clonal individuals as well as the blooming properties of male and female flowers at the same time. This study could pro-vide theoretical reference for working out the production plan of improved varieties and other management measures in seed orchard of Larix principis-rupprechtii.展开更多
Relationships between stem growth and climatic and edaphic factors,notably air temperatures and soil moisture for different slopes,are not completely understood.Stem radial variations were monitored at the bottom and ...Relationships between stem growth and climatic and edaphic factors,notably air temperatures and soil moisture for different slopes,are not completely understood.Stem radial variations were monitored at the bottom and top slope positions in a Larix principis-rupprechtii plantation during the 2017 and 2018 growing seasons.Total precipitation during the growing season in 2017 and 2018 was 566 mm and 728 mm,respectively.Stem contractions typically occurred after mid-morning followed by swelling in the late afternoon in both plots,reflecting the diurnal cycle of water uptake and loss.Trees at the two locations showed the same growth initiation(mid-May)because of the small differences in air and soil temperatures.There were no significant differences in cumulative stem radial growth between the bottom plot(1.57±0.34 mm)and the top plot(1.55±0.26 mm)in 2018.However,in 2017,the main growth period of the bottom plot ceased 17 days earlier than in the top plot,while cumulative seasonal growth of the bottom plot(1.08±0.25 mm)was significantly less than the top plot(1.54±0.43 mm).Maximum daily stem shrinkage was positively correlated with air and soil temperatures,solar radiation,vapor pressure deficits,and negatively correlated with volumetric soil moisture content.The maximum daily shrinkage reflected transpiration rates as affected by environmental factors.Daily radial stem increment was correlated with precipitation and volumetric soil moisture in both years,but with air temperatures only in 2017.The seasonal growth of L.principis-rupprechtii Mayr thus shows interannual dynamics,while precipitation constitutes a key driving factor.展开更多
Larix principis-rupprechtii Mayr is a dominant species in coniferous forests of North China.However,early selection of L.principis-rupprechtii for growth traits is poorly characterised.To explore the optimal selection...Larix principis-rupprechtii Mayr is a dominant species in coniferous forests of North China.However,early selection of L.principis-rupprechtii for growth traits is poorly characterised.To explore the optimal selection age for this species,heights(HT)and diameters at breast(DBH)of 40 half-sib families were measured at ages 3,12,22,and 28 years in a progeny test population established in the town of Kangjiahui,Shanxi Province.Age trends in heritability,age–age genetic correlations,and early selection efficiency for height and DBH were analysed.The individual heritability of these growth traits varied over time,and maximized at different ages(0.55 at age 12 for HT and 0.48 at age 28 for DBH).The age–age genetic correlations were always positive,and the majority were high(0.790–0.953)between the juvenile and mature ages for HT and DBH.For the same pairs of measurements,HT demonstrated higher age–age genetic correlations than DBH,and both age–age genetic correlation data sets were described well by the linear relationship with the logarithm of the age ratio(r2[0.90).The regression slope for DBH was lower than that for HT.Based on the early selection efficiency estimates,the optimal selection age could be as early as age 6 for DBH and 8–9 years for HT.The results of this study provide information that can be used to assist early selection practices in L.principis-rupprechtii improvement programs in Shanxi Province.展开更多
Horizontal and veftical distributions of fine root abundance (mass per unit ground surface arca) were investi-gated in a densely planted larch - ash -stripe - mixed tbrest on dark broxvn forest soil in northeast China...Horizontal and veftical distributions of fine root abundance (mass per unit ground surface arca) were investi-gated in a densely planted larch - ash -stripe - mixed tbrest on dark broxvn forest soil in northeast China. There was evidencefor some degree of below-ground niche partitioning (or differentiation) betmpen the two species in hoth the horizontal andvertical directions. The ash fine roots largely penetrated into the larch belt (larch Sub-community) in surttrce soil (0-20 cm),indicating a possible inductive effect of larch ecological field on ash roots, while the penetration of larch tine roots into ashbelt (ash sub-community) was much restricted- which reflected a negative influence of ash ccological tield on Iarch roots. Inthe vertical direction of marginal soil the ash tine roots were mainly distributed in topsoil with a vertical gradient similar tothat as in the internal ash sub-community. but the larch fine roots were relatively compelled to deepcr soil layers by the com-petition (or exclusion) of marginal ash trees. All the dit1brences or complemcntarity wcre considered to the result of inter-specitic competition, which was important to the coexistence of the two tbrest species and the sustainability of mixed planta-tion.展开更多
Leaf area index(LAI) is a key factor that determines a forest ecosystem's net primary production and energy exchange between the atmosphere and land surfaces.LAI can be measured in many ways, but there has been li...Leaf area index(LAI) is a key factor that determines a forest ecosystem's net primary production and energy exchange between the atmosphere and land surfaces.LAI can be measured in many ways, but there has been little research to compare LAI estimated by different methods. In this study, we compared the LAI results from two different approaches, i.e., the dimidiate pixel model(DPM) and an empirical statistic model(ESM) using ZY-3 high-accuracy satellite images validated by field data. We explored the relationship of LAI of Larix principis-rupprechtii Mayr plantations with topographic conditions. The results show that DPM improves the simulation of LAI(r = 0.86,RMSE = 0.57) compared with ESM(r = 0.62, RMSE =0.79). We further concluded that elevation and slope significantly affect the distribution of LAI. The maximum peak of LAI appeared at an aspect of east and southeast at an elevation of 1700–2000 m. Our results suggest that ZY-3 can satisfy the needs of quantitative monitoring of leaf area indices in small-scale catchment areas. DPM provides a simple and accurate method to obtain forest vegetation parameters in the case of non-ground measurement points.展开更多
Soil samples were collected from the surface soil of three kinds of plantations mixed plantation of Dahurian larch and Manchurian ash, pure ash plantation and pure larch plantation, at Maoershan Forest experiment cent...Soil samples were collected from the surface soil of three kinds of plantations mixed plantation of Dahurian larch and Manchurian ash, pure ash plantation and pure larch plantation, at Maoershan Forest experiment center in Heilongjiang Province. The result of measuring and analysis showed that the number of the bacteria was the most and their effective scope and active strength were the most remarkable comparing with various kinds of soil microorganisms. Actinomyces were the second most and the fungi were the least in number Numbers of the soil microorganisms in mixed plantation had a more number than that in the pure ash plantation. The number and species of microorganisms were different among different soil layers. For the same kind of soil, the number and species of the microorganisms in the surface layer were the most, and declined wth the increase of the soil depth. The seasonal change of the number of the soil microorganisms in pure or mixed pIantations was also remarkable. The number of the microorganisms increases obviously with the increase of the atmospheric temperature. The total number of microorganisms in Jan. was 30.0%-40.2% of the number in Sep. Both the vertical distribution and seasonal change being concerned, the number of the soil microorganisms in mixed plantation was on a higher level than that in pure plantation.展开更多
Plot sampling was conducted in the second cultivation areas ofLarix olgensis in Heilongjiang Province, China. By analyzing plot investigation data, wood properties, and kraft pulps of 840 plots and 248 sample trees in...Plot sampling was conducted in the second cultivation areas ofLarix olgensis in Heilongjiang Province, China. By analyzing plot investigation data, wood properties, and kraft pulps of 840 plots and 248 sample trees in industrial plantations of L. olgensis on different sites, we examined the growth process of L. olgensis industrial plantation with suitable structure, the wood fiber feature, chemical composition, physical performance and pulp characteristics. It is suggested that site conditions have major effects on plantation growth, fiber contains, fiber length, rate between fiber length and fiber width, pulping rate and pulp physics intensity. The best site for L. olgensis industrial plantation growth is site class Ⅰ and site class Ⅱ, which are on lower locations. Site condition has an obvious influence on the wood characteristics. Within the range of site conditions and stand densities studied, the worse the site condition, the less the fiber contains, the shorter the fiber length, and the more the 1% NaOH extraction. This kind of relationship becomes more obvious as stand age increases. However, the influence of site condition on pulping rate and pulp physics intensity is not obvious. The result provides theoretical base for cultivation of industrial fiber plantation of L. olgensis.展开更多
In this study,three different ages of Larix principis-rupprechtii forests in 5,10 and 20 years were selected as the research objects,and the changes in soil nutrient and soil enzyme activities in different growth stag...In this study,three different ages of Larix principis-rupprechtii forests in 5,10 and 20 years were selected as the research objects,and the changes in soil nutrient and soil enzyme activities in different growth stages were analyzed. The results showed that the contents of organic matter and available phosphorus in the soil of different growth stages showed a significant downward trend with the increase of soil depth.For different forest ages in the same soil layer,the soil available phosphorus content declined with the increase of the forest age. The organic matter content of 40-60 cm in 20 years of forest age was the lowest in July,which was 4. 17 g/kg,significantly lower than that in other soil layers. Besides,the soil available phosphorus content of 5 years of forest age reached the maximum in July,with an average of 4. 44 mg/kg,which was higher than the available phosphorus content in soil in May and September,but the difference between the three months was not significant. The changes in ammonium nitrogen and nitrate nitrogen content in soil with different forest ages were consistent with the changes in the soil depth,showing a downward trend. In the new leaf stage,the ammonium nitrogen content of the L. principis-rupprechtii forest land in5 years of age at 20-40 and 40-60 cm of the soil depth was 13. 47 and 9. 09 mg/kg,respectively,which was 46. 9% and 64. 2% lower than that at 0-20 cm( 25. 36 mg/kg) of the soil depth. The soil nitrate nitrogen content of 20 years of forest age was 19. 24 mg/kg,which was25. 8% lower than that of 10 years of forest age,showing significant difference( P < 0. 05). In addition,with the increase of the age of L. principis-rupprechtii,soil catalase( CAT),alkaline phosphatase( ALP) and urease( Ure) decreased,and the decline of ALP was slow,while CAT and Ure decreased significantly. In summary,it is concluded that the soil fertility of forest land declined with the increase of forest age on the basis of change trend of soil nutrient and soil enzyme activity in the surveyed forest age.展开更多
Background:Stem CO_(2) efflux(E_(S))plays a critical role in the carbon budget of forest ecosystems.Thinning is a core practice for sustainable management of plantations.It is therefore necessary and urgent to study t...Background:Stem CO_(2) efflux(E_(S))plays a critical role in the carbon budget of forest ecosystems.Thinning is a core practice for sustainable management of plantations.It is therefore necessary and urgent to study the effect and mechanism of thinning intensity(TI)on E_(S).Methods:In this study,five TIs were applied in Larix principis-rupprechtii Mayr 21-,25-,and 41-year-old stands in North China in 2010.Portable infrared gas analyzer(Li-8100 A)was used to measure ES and its association with environmental factors at monthly intervals from May to October in 2013 to 2015.In addition,nutrients,wood structure and nonstructural carbon(NSC)data were measured in August 2016.Results:The results show that ES increased with increasing TI.The maximum ES values occurred at a TI of 35%(3.29,4.57 and 2.98μmol·m^(-2)·s^(-1))and were 1.54-,1.94-and 2.89-fold greater than the minimum E_(S) value in the CK stands(2.14,2.35 and 1.03μmol·m^(-2)·s^(-1))in July for the 21-,25-and 41-year-old forests,respectively.The E_(S) of the trees in low-density stands was more sensitive to temperature than that of the trees in high-density stands.Soluble sugars(SS)and temperature are the main factors affecting ES.When the stand density is low enough as 41-year-old L.principis-rupprechtii forests with TI 35%,bark thickness(BT)and humidity should be considered in addition to air temperature(T_(a)),wood temperature(T_(w)),sapwood width(SW),nitrogen concentration(N)and SS in the evaluation of ES.If a change in stand density is ignored,the CO_(2) released from individual 21-,25-and 41-year-old trees could be underestimated by 168.89%,101.94% and 200.49%,respectively.CO_(2) release was estimated based on the stem equation in combination with the factors influencing ES for reference.Conclusions:We suggest that it is not sufficient to conventional models which quantify ES only by temperature and that incorporating the associated drivers(e.g.density,SS,SW and N)based on stand density into conventional models can improve the accuracy of ES estimates.展开更多
Prince Rupprecht’s larch(Larix principis-rupprechtii Mayr.),a deciduous conifer,widely grows in middle and high elevations of Northern China.Its natural distribution has sharply decreased and has become fragmented,wh...Prince Rupprecht’s larch(Larix principis-rupprechtii Mayr.),a deciduous conifer,widely grows in middle and high elevations of Northern China.Its natural distribution has sharply decreased and has become fragmented,which may have resulted in the loss of genetic variation.In this study,ten natural populations across the entire range of this species were analyzed using amplifi ed fragment length polymorphism markers.A total of 309 loci were detected from 225 individuals of these populations,of which 261(84.5%)were polymorphic.At the species level,the genetic diversity was high(average of the Nei’s genetic diversity H e=0.2602,and Shannon’s information index I=0.3967).The results of molecular variance analysis showed that 90.71%of the genetic diversity occurred within populations.The genetic diff erentiation among populations was moderate as a whole(F ST=0.0929,G ST=0.1510),which is consistent with the moderate level of gene fl ow among populations(N m=2.8116).Based on the unweighted pair group method with arithmetic mean and STRU CTU RE analysis,these populations were grouped into three genetically distinct clusters.The degree of inter-population diff erentiation(G ST=0.1338)for the south group was larger than that for the north group(G ST=0.0915).There was a signifi cant correlation between genetic distance and geographic distance across the species range(r=0.316,P<0.05).Genetic diversity was signifi-cantly associated with longitude but not elevation or climatic factors.The populations with high genetic diversity from each cluster are therefore recommended for future conservation and management of this species.展开更多
Several equations were selected using nonlinear regression analysis for setting up growth and yield modehe of Dahurian larch (Laris gmelinii Rupr.) plantations. Data of 405 stem analysis trees were collected from 336 ...Several equations were selected using nonlinear regression analysis for setting up growth and yield modehe of Dahurian larch (Laris gmelinii Rupr.) plantations. Data of 405 stem analysis trees were collected from 336 temporary plots throughout the Daxing’an Mountains. Results showed that the Richards equation was the best model for estimating tree height, stand mean height and stand dominant height by age, the Power equation was the fdiest model for predicting tree volume by DBH and tree height, and the Logarithmic stand vofume equation was good for predicting stand volume from age, mean height. basal area and other stand variables. These models can be used to construct volume tabIes, site index table and other forestry tables for Dahurian ghantations.展开更多
Community properties of dahurian larch (Larix gmelini) plantations at the Experimental Station of Laoshan were studied. Results show that under the different distributions of the combines of tree layer of dahurian lar...Community properties of dahurian larch (Larix gmelini) plantations at the Experimental Station of Laoshan were studied. Results show that under the different distributions of the combines of tree layer of dahurian larch plantations, stands have different community structure characteristics and biological productivities. From the result of the analysis of variance, it shows that there is the obvious difference among biomass of understory, herb layer and litter for four structure tpps.展开更多
The biornass and productivity for dahurian larch (Larix gmelini) plantations of four structure types were measured. The biomass of trees, understory' herb and litter was surveyed using methods of average standard ...The biornass and productivity for dahurian larch (Larix gmelini) plantations of four structure types were measured. The biomass of trees, understory' herb and litter was surveyed using methods of average standard tree and subsample yield respectively.The estimated total mean biomass for dahurian larch plantation communities of four structure types was 159. 445 t/hm2 and the net productivity was 12. 237 t/hm2. a.展开更多
Research has indicated that simple forest eco-system composition,structure and diversity have uncompli-cated community relationships and insufficient pest control capabilities.To investigate changing characteristics o...Research has indicated that simple forest eco-system composition,structure and diversity have uncompli-cated community relationships and insufficient pest control capabilities.To investigate changing characteristics of plant and insect communities in under pest outbreaks in Larix principis-rupprechtii plantations,the research areas were defined as mature(48–50 years)and young(24–29 years)infested stands along with healthy stands.The results show a reduction in the complexity and diversity of plant communi-ties and herbaceous plant guilds(polycultures of beneficial plants)and the complexity and dominance of insect com-munities,especially natural insect enemies.The results also show the relative simplicity of the main factors of commu-nity change and development that represent the characteris-tics of pest outbreaks in L.principis-rupprechtii plantations.The complexity and diversity of plant communities,particu-larly herbaceous plant guilds play a fundamental role in the regulation and development in forest ecosystems.展开更多
基金the National Natural Science Foundation of China(Nos.42161144008U21A2005+3 种基金U20A2085)the National Key Research and Development Program of China(2022YFF08018042022YFF08018032022YFF1300404)。
文摘Extreme climate has increasingly led to negative impacts on forest ecosystems globally,especially in semiarid areas where forest ecosystems are more vulnerable.However,it is poorly understood how tree growth is affected by different drought events.In 2006–2009,the larch plantations in the semiarid areas of Northwest China were negatively affected by four consecutive dry years,which was a very rare phenomenon that may occur frequently under future climate warming.In this study,we analyzed the effect of these consecutive dry years on tree growth based on the data of the tree rings in the dominant layer of the forest canopy on a larch plantation.We found that the tree-ring width index(RWI)in dry years was lower than that in normal years,and it experienced a rapidly decreasing trend from 2006 to 2009(slope=-0.139 year^(-1),r=-0.94)due to water supply deficits in those dry years.Drought induced legacy effects of tree growth reduction,and consecutive dry years corresponded with greater growth reductions and legacy effects.Growth reductions and legacy effects were significantly stronger in the third and fourth consecutive dry years than that of single dry year(p<0.05),which might have been due to the cumulative stress caused by consecutive dry years.Our results showed that larch trees experienced greater tree growth reduction due to consecutive dry years and their legacy effect,and the trees had lower recovery rates after consecutive dry years.Our results highlight that consecutive dry years pose a new threat to plantations under climate warming,and thus,the effect of climate extremes on tree growth should be considered in growth models in semiarid areas.
基金supported by the Key Research and Development Program of Hebei Province[20326333D]Science and Technology Promotion Demonstration Project of forestry and grassland in Hebei Province[Hebei TG[2019]001]Science and Technology Promotion Demonstration Project of forestry and grassland in Hebei Province[Hebei TG[2020]013]。
文摘Larix gmelinii var.principis-rupprechtii(Mayr.)Pilger is an important native tree species in North China with advantages of fast growth,straight trunk,and good wood properties.The multi-year and multi-site breeding research of families of the species has not been reported previously.Based on diameter at breast height(DBH),height and volume of 25 families on four experimental sites,we calculated variance components,genetic parameters,juvenile and mature trait correlations and made genotype main effect plus genotype×environment interaction effect(GGE)biplot based on the breeding values estimated using the method of best linear unbiased prediction(BLUP).Compared with height,DBH and volume had higher heritability and larger variation coefficients,making them the more suitable traits for family selection and evaluation.Based on these,GGE biplots containing 20 combinations of site×age were drawn using data at 13 to 17 years when the interactions between family and location were strong.Test sites classifications based on DBH,and volume were inconsistent,with two categories for DBH and one for volume.The Guyuan site was the most suitable with strong discriminating ability,high representativeness and stability among tree ages.Integrating the ranking results of DBH and volume,families 66,76,82 and 111 were high-yielding and stable,families 78 and96 were high-yielding with above average stability,families72 and 79 were high-yielding with below average stability,whereas stability of family 100 was inconsistent between DBH and volume.Early selection based on DBH was convenient and reliable,and can be made at seven years.This study provides support for the selection of Larix gmelinii var.principis-rupprechtii families in Hebei province and an example for the application of stem analysis data from multiple sites in tree breeding.
基金supported by the National Key Research and Development Program of China(2017YFD0600401)the Fundamental Research Funds for the Central Universities(2572019CP08)
文摘Korean larch(Larix olgensis)is one of the main tree species for aff orestation and timber production in northeast China.However,its timber quality and growth ability are largely infl uenced by crown size,structure and shape.The majority of crown models are static models based on tree size and stand characteristics from temporary sample plots,but crown dynamic models has seldom been constructed.Therefore,this study aimed to develop height to crown base(HCB)and crown length(CL)dynamic models using the branch mortality technique for a Korean larch plantation.The nonlinear mixed-eff ects model with random eff ects,variance functions and correlation structures,was used to build HCB and CL dynamic models.The data were obtained from 95 sample trees of 19 plots in Meng JiaGang forest farm in Northeast China.The results showed that HCB progressively increases as tree age,tree height growth(HT growth)and diameter at breast height growth(DBH growth).The CL was increased with tree age in 20 years ago,and subsequently stabilized.HT growth,DBH growth stand basal area(BAS)and crown competition factor(CCF)signifi cantly infl uenced HCB and CL.The HCB was positively correlated with BAS,HT growth and DBH growth,but negatively correlated with CCF.The CL was positively correlated with BAS and CCF,but negatively correlated with DBH growth.Model fi tting and validation confi rmed that the mixed-eff ects model considering the stand and tree level random eff ects was accurate and reliable for predicting the HCB and CL dynamics.However,the models involving adding variance functions and time series correlation structure could not completely remove heterogeneity and autocorrelation,and the fi tting precision of the models was reduced.Therefore,from the point of view of application,we should take care to avoid setting up over-complex models.The HCB and CL dynamic models in our study may also be incorporated into stand growth and yield model systems in China.
基金This study was supported by “the 100-Young-Research Project” of Chinese Academy of Sciences and National Natural Science Foundation of China (30371149)
文摘In order to understand the effects of thinning on microsite conditions and natural regeneration in the larch plantation, thinning experiment was conducted in a 40-year-old Larix olgensis plantation in Qingyuan County in eastern Liaoning Province, China in 2003-2004 Five thinning treatments (0%, 10.2%, 19.8%, 29.7% and 40.3% thinned) were designed on the same site. After thinning, canopy openness and the microsite conditions such as photosynthetic photon flux density (PPFD), soil moisture content, and soil temperature were measured in one growing season. Meanwhile, the investigation of natural regeneration was conducted at the end of the growing season. The results showed that the canopy openness increased with the increase of thinning intensities. PPFD and soil temperature and soil moisture content in different soil layers were positively relative with canopy openness after thinning. The richness of regenerating tree species did not significantly increase (p=0.30) after one growing season since thinning, but the regeneration density and frequency of tree species increased significantly (p〈0.05). In addition, the number of regenerating tree species increased, and the increment was correlated with the characteristics of iudividual tree species. The increasing percentage of regenerating seedlings of the shade-intolerant tree species was more than that of shade-tolerant tree species. Among the investigated regeneration species, the biggest response of seedling emergency to the canopy openness was Phellodendron amurense. This paper confirmed the following conclusions: after thinning, the variety of regenerating tree species was correlative with the characteristics of regenerating tree species, and the distribution of unthinned trees and the site conditions in the investigated larch plantation were the additional factors influencing, the regeneration.
基金financially supported by the National Sci-Tech Support Plan of China(Grant No.2015BAD14B05)
文摘Because growth ring data have temporal features, time series analysis can be used to simulate and reveal changes in the life of a tree and contribute to plantation management. In this study, the autoregressive(AR) and moving average modeling method was used to simulate the time series for growth ring density in a larch plantation with different initial planting densities. We adopted the Box–Jenkins method for the modeling, which was initially based on an intuitive analysis of sequence graphs followed by the augmented Dickey–Fuller stationarity test. The order p and q of the ARMA(p, q) model was determined based on the autocorrelation and partial correlation coefficient figure truncated on the respective order.Through the residual judgment, the model AR(2) was only fitted to the larch growth ring density series for the plantation with the 1.5 9 2.0 m^2 initial planting density.Because the residuals series for the other three series was not shown as a white noise sequence, the modeling was rerun. Larch wood from the initial planting density of2.0 9 2.0 m^2 was modeled by ARMA(2, 1), and ARMA((1, 5), 3) fitted to the 2.5 9 2.5 m^2 initial planting density,and the 3.0 9 3.0 m^2 was modeled by AR(1, 2, 5).Although the ARMA modeling can simulate the change in growth ring density, data for the different growth ring time series were described by different models. Thus, time series modeling can be suitable for growth ring data analysis, revealing the time domain and frequency domain of growth ring data.
文摘Through 5 years of phenological observations on Larix principis-rupprechtii Mayr. in primary seed orchard and studies on population and individuals of clones, the annual periodic phenological laws were revealed and the annual phe-nological periodic table was drawn up. The correlation between various phenophases, the air temperature and active accumu-lated temperature were analyzed and expounded. The authors also analyzed the similarities and differences of phenophases among clonal individuals as well as the blooming properties of male and female flowers at the same time. This study could pro-vide theoretical reference for working out the production plan of improved varieties and other management measures in seed orchard of Larix principis-rupprechtii.
基金supported by the Ningxia Hui Autonomous Region(QCYL-2018-12)the National Key Research and Development Program of China(2016YFC0501603,2017YFC0504602)the National Natural Science Foundation of China(41671025).
文摘Relationships between stem growth and climatic and edaphic factors,notably air temperatures and soil moisture for different slopes,are not completely understood.Stem radial variations were monitored at the bottom and top slope positions in a Larix principis-rupprechtii plantation during the 2017 and 2018 growing seasons.Total precipitation during the growing season in 2017 and 2018 was 566 mm and 728 mm,respectively.Stem contractions typically occurred after mid-morning followed by swelling in the late afternoon in both plots,reflecting the diurnal cycle of water uptake and loss.Trees at the two locations showed the same growth initiation(mid-May)because of the small differences in air and soil temperatures.There were no significant differences in cumulative stem radial growth between the bottom plot(1.57±0.34 mm)and the top plot(1.55±0.26 mm)in 2018.However,in 2017,the main growth period of the bottom plot ceased 17 days earlier than in the top plot,while cumulative seasonal growth of the bottom plot(1.08±0.25 mm)was significantly less than the top plot(1.54±0.43 mm).Maximum daily stem shrinkage was positively correlated with air and soil temperatures,solar radiation,vapor pressure deficits,and negatively correlated with volumetric soil moisture content.The maximum daily shrinkage reflected transpiration rates as affected by environmental factors.Daily radial stem increment was correlated with precipitation and volumetric soil moisture in both years,but with air temperatures only in 2017.The seasonal growth of L.principis-rupprechtii Mayr thus shows interannual dynamics,while precipitation constitutes a key driving factor.
基金supported by National Key R&D Program of China(2017YFD0600404-1)Medium and Long Scientific Research Project for Young Teachers in Beijing Forestry University(2015ZCQ-SW-02)+2 种基金the Project of National Natural Science Foundation of China(31370658)Program for Changjiang Scholars and Innovative Research Team in University(IRT13047)the Project of Beijing Gardening and Greening Bureau(CEG-2016-01)
文摘Larix principis-rupprechtii Mayr is a dominant species in coniferous forests of North China.However,early selection of L.principis-rupprechtii for growth traits is poorly characterised.To explore the optimal selection age for this species,heights(HT)and diameters at breast(DBH)of 40 half-sib families were measured at ages 3,12,22,and 28 years in a progeny test population established in the town of Kangjiahui,Shanxi Province.Age trends in heritability,age–age genetic correlations,and early selection efficiency for height and DBH were analysed.The individual heritability of these growth traits varied over time,and maximized at different ages(0.55 at age 12 for HT and 0.48 at age 28 for DBH).The age–age genetic correlations were always positive,and the majority were high(0.790–0.953)between the juvenile and mature ages for HT and DBH.For the same pairs of measurements,HT demonstrated higher age–age genetic correlations than DBH,and both age–age genetic correlation data sets were described well by the linear relationship with the logarithm of the age ratio(r2[0.90).The regression slope for DBH was lower than that for HT.Based on the early selection efficiency estimates,the optimal selection age could be as early as age 6 for DBH and 8–9 years for HT.The results of this study provide information that can be used to assist early selection practices in L.principis-rupprechtii improvement programs in Shanxi Province.
文摘Horizontal and veftical distributions of fine root abundance (mass per unit ground surface arca) were investi-gated in a densely planted larch - ash -stripe - mixed tbrest on dark broxvn forest soil in northeast China. There was evidencefor some degree of below-ground niche partitioning (or differentiation) betmpen the two species in hoth the horizontal andvertical directions. The ash fine roots largely penetrated into the larch belt (larch Sub-community) in surttrce soil (0-20 cm),indicating a possible inductive effect of larch ecological field on ash roots, while the penetration of larch tine roots into ashbelt (ash sub-community) was much restricted- which reflected a negative influence of ash ccological tield on Iarch roots. Inthe vertical direction of marginal soil the ash tine roots were mainly distributed in topsoil with a vertical gradient similar tothat as in the internal ash sub-community. but the larch fine roots were relatively compelled to deepcr soil layers by the com-petition (or exclusion) of marginal ash trees. All the dit1brences or complemcntarity wcre considered to the result of inter-specitic competition, which was important to the coexistence of the two tbrest species and the sustainability of mixed planta-tion.
基金supported by the National Forestry Public Welfare Professional Scientific Research Project(No.201404213)the National Key Research and Development Program of China(No.2016YFD0600205)
文摘Leaf area index(LAI) is a key factor that determines a forest ecosystem's net primary production and energy exchange between the atmosphere and land surfaces.LAI can be measured in many ways, but there has been little research to compare LAI estimated by different methods. In this study, we compared the LAI results from two different approaches, i.e., the dimidiate pixel model(DPM) and an empirical statistic model(ESM) using ZY-3 high-accuracy satellite images validated by field data. We explored the relationship of LAI of Larix principis-rupprechtii Mayr plantations with topographic conditions. The results show that DPM improves the simulation of LAI(r = 0.86,RMSE = 0.57) compared with ESM(r = 0.62, RMSE =0.79). We further concluded that elevation and slope significantly affect the distribution of LAI. The maximum peak of LAI appeared at an aspect of east and southeast at an elevation of 1700–2000 m. Our results suggest that ZY-3 can satisfy the needs of quantitative monitoring of leaf area indices in small-scale catchment areas. DPM provides a simple and accurate method to obtain forest vegetation parameters in the case of non-ground measurement points.
文摘Soil samples were collected from the surface soil of three kinds of plantations mixed plantation of Dahurian larch and Manchurian ash, pure ash plantation and pure larch plantation, at Maoershan Forest experiment center in Heilongjiang Province. The result of measuring and analysis showed that the number of the bacteria was the most and their effective scope and active strength were the most remarkable comparing with various kinds of soil microorganisms. Actinomyces were the second most and the fungi were the least in number Numbers of the soil microorganisms in mixed plantation had a more number than that in the pure ash plantation. The number and species of microorganisms were different among different soil layers. For the same kind of soil, the number and species of the microorganisms in the surface layer were the most, and declined wth the increase of the soil depth. The seasonal change of the number of the soil microorganisms in pure or mixed pIantations was also remarkable. The number of the microorganisms increases obviously with the increase of the atmospheric temperature. The total number of microorganisms in Jan. was 30.0%-40.2% of the number in Sep. Both the vertical distribution and seasonal change being concerned, the number of the soil microorganisms in mixed plantation was on a higher level than that in pure plantation.
基金Foundation project: This paper was supported by China Postdoctoral Science Foundation (2005037637), Heilongjiang Postdoctoral Science Foundation (LBH-Z05031) and Project of State Forestry Administration, China (2006-37).
文摘Plot sampling was conducted in the second cultivation areas ofLarix olgensis in Heilongjiang Province, China. By analyzing plot investigation data, wood properties, and kraft pulps of 840 plots and 248 sample trees in industrial plantations of L. olgensis on different sites, we examined the growth process of L. olgensis industrial plantation with suitable structure, the wood fiber feature, chemical composition, physical performance and pulp characteristics. It is suggested that site conditions have major effects on plantation growth, fiber contains, fiber length, rate between fiber length and fiber width, pulping rate and pulp physics intensity. The best site for L. olgensis industrial plantation growth is site class Ⅰ and site class Ⅱ, which are on lower locations. Site condition has an obvious influence on the wood characteristics. Within the range of site conditions and stand densities studied, the worse the site condition, the less the fiber contains, the shorter the fiber length, and the more the 1% NaOH extraction. This kind of relationship becomes more obvious as stand age increases. However, the influence of site condition on pulping rate and pulp physics intensity is not obvious. The result provides theoretical base for cultivation of industrial fiber plantation of L. olgensis.
文摘In this study,three different ages of Larix principis-rupprechtii forests in 5,10 and 20 years were selected as the research objects,and the changes in soil nutrient and soil enzyme activities in different growth stages were analyzed. The results showed that the contents of organic matter and available phosphorus in the soil of different growth stages showed a significant downward trend with the increase of soil depth.For different forest ages in the same soil layer,the soil available phosphorus content declined with the increase of the forest age. The organic matter content of 40-60 cm in 20 years of forest age was the lowest in July,which was 4. 17 g/kg,significantly lower than that in other soil layers. Besides,the soil available phosphorus content of 5 years of forest age reached the maximum in July,with an average of 4. 44 mg/kg,which was higher than the available phosphorus content in soil in May and September,but the difference between the three months was not significant. The changes in ammonium nitrogen and nitrate nitrogen content in soil with different forest ages were consistent with the changes in the soil depth,showing a downward trend. In the new leaf stage,the ammonium nitrogen content of the L. principis-rupprechtii forest land in5 years of age at 20-40 and 40-60 cm of the soil depth was 13. 47 and 9. 09 mg/kg,respectively,which was 46. 9% and 64. 2% lower than that at 0-20 cm( 25. 36 mg/kg) of the soil depth. The soil nitrate nitrogen content of 20 years of forest age was 19. 24 mg/kg,which was25. 8% lower than that of 10 years of forest age,showing significant difference( P < 0. 05). In addition,with the increase of the age of L. principis-rupprechtii,soil catalase( CAT),alkaline phosphatase( ALP) and urease( Ure) decreased,and the decline of ALP was slow,while CAT and Ure decreased significantly. In summary,it is concluded that the soil fertility of forest land declined with the increase of forest age on the basis of change trend of soil nutrient and soil enzyme activity in the surveyed forest age.
基金funding from National Natural Science Foundation of China(No.31870387),China Scholarship Council.
文摘Background:Stem CO_(2) efflux(E_(S))plays a critical role in the carbon budget of forest ecosystems.Thinning is a core practice for sustainable management of plantations.It is therefore necessary and urgent to study the effect and mechanism of thinning intensity(TI)on E_(S).Methods:In this study,five TIs were applied in Larix principis-rupprechtii Mayr 21-,25-,and 41-year-old stands in North China in 2010.Portable infrared gas analyzer(Li-8100 A)was used to measure ES and its association with environmental factors at monthly intervals from May to October in 2013 to 2015.In addition,nutrients,wood structure and nonstructural carbon(NSC)data were measured in August 2016.Results:The results show that ES increased with increasing TI.The maximum ES values occurred at a TI of 35%(3.29,4.57 and 2.98μmol·m^(-2)·s^(-1))and were 1.54-,1.94-and 2.89-fold greater than the minimum E_(S) value in the CK stands(2.14,2.35 and 1.03μmol·m^(-2)·s^(-1))in July for the 21-,25-and 41-year-old forests,respectively.The E_(S) of the trees in low-density stands was more sensitive to temperature than that of the trees in high-density stands.Soluble sugars(SS)and temperature are the main factors affecting ES.When the stand density is low enough as 41-year-old L.principis-rupprechtii forests with TI 35%,bark thickness(BT)and humidity should be considered in addition to air temperature(T_(a)),wood temperature(T_(w)),sapwood width(SW),nitrogen concentration(N)and SS in the evaluation of ES.If a change in stand density is ignored,the CO_(2) released from individual 21-,25-and 41-year-old trees could be underestimated by 168.89%,101.94% and 200.49%,respectively.CO_(2) release was estimated based on the stem equation in combination with the factors influencing ES for reference.Conclusions:We suggest that it is not sufficient to conventional models which quantify ES only by temperature and that incorporating the associated drivers(e.g.density,SS,SW and N)based on stand density into conventional models can improve the accuracy of ES estimates.
基金the Natural Science Foundation of Shanxi Province,China(2010011041-1)the National Natural Science Foundation of China(41271531).
文摘Prince Rupprecht’s larch(Larix principis-rupprechtii Mayr.),a deciduous conifer,widely grows in middle and high elevations of Northern China.Its natural distribution has sharply decreased and has become fragmented,which may have resulted in the loss of genetic variation.In this study,ten natural populations across the entire range of this species were analyzed using amplifi ed fragment length polymorphism markers.A total of 309 loci were detected from 225 individuals of these populations,of which 261(84.5%)were polymorphic.At the species level,the genetic diversity was high(average of the Nei’s genetic diversity H e=0.2602,and Shannon’s information index I=0.3967).The results of molecular variance analysis showed that 90.71%of the genetic diversity occurred within populations.The genetic diff erentiation among populations was moderate as a whole(F ST=0.0929,G ST=0.1510),which is consistent with the moderate level of gene fl ow among populations(N m=2.8116).Based on the unweighted pair group method with arithmetic mean and STRU CTU RE analysis,these populations were grouped into three genetically distinct clusters.The degree of inter-population diff erentiation(G ST=0.1338)for the south group was larger than that for the north group(G ST=0.0915).There was a signifi cant correlation between genetic distance and geographic distance across the species range(r=0.316,P<0.05).Genetic diversity was signifi-cantly associated with longitude but not elevation or climatic factors.The populations with high genetic diversity from each cluster are therefore recommended for future conservation and management of this species.
文摘Several equations were selected using nonlinear regression analysis for setting up growth and yield modehe of Dahurian larch (Laris gmelinii Rupr.) plantations. Data of 405 stem analysis trees were collected from 336 temporary plots throughout the Daxing’an Mountains. Results showed that the Richards equation was the best model for estimating tree height, stand mean height and stand dominant height by age, the Power equation was the fdiest model for predicting tree volume by DBH and tree height, and the Logarithmic stand vofume equation was good for predicting stand volume from age, mean height. basal area and other stand variables. These models can be used to construct volume tabIes, site index table and other forestry tables for Dahurian ghantations.
文摘Community properties of dahurian larch (Larix gmelini) plantations at the Experimental Station of Laoshan were studied. Results show that under the different distributions of the combines of tree layer of dahurian larch plantations, stands have different community structure characteristics and biological productivities. From the result of the analysis of variance, it shows that there is the obvious difference among biomass of understory, herb layer and litter for four structure tpps.
文摘The biornass and productivity for dahurian larch (Larix gmelini) plantations of four structure types were measured. The biomass of trees, understory' herb and litter was surveyed using methods of average standard tree and subsample yield respectively.The estimated total mean biomass for dahurian larch plantation communities of four structure types was 159. 445 t/hm2 and the net productivity was 12. 237 t/hm2. a.
文摘Research has indicated that simple forest eco-system composition,structure and diversity have uncompli-cated community relationships and insufficient pest control capabilities.To investigate changing characteristics of plant and insect communities in under pest outbreaks in Larix principis-rupprechtii plantations,the research areas were defined as mature(48–50 years)and young(24–29 years)infested stands along with healthy stands.The results show a reduction in the complexity and diversity of plant communi-ties and herbaceous plant guilds(polycultures of beneficial plants)and the complexity and dominance of insect com-munities,especially natural insect enemies.The results also show the relative simplicity of the main factors of commu-nity change and development that represent the characteris-tics of pest outbreaks in L.principis-rupprechtii plantations.The complexity and diversity of plant communities,particu-larly herbaceous plant guilds play a fundamental role in the regulation and development in forest ecosystems.