Hopf bifurcation and phase synchronization in memristor-coupled Hindmarsh–Rose and FitzHugh–Nagumo neurons with two time delays

A memristor-coupled heterogenous neural network consisting of two-dimensional(2D)FitzHugh–Nagumo(FHN)and Hindmarsh–Rose(HR)neurons with two time delays is established.Taking the time delays as the control parameters,the existence of Hopf bifurcation near the stable equilibrium point in four cases is derived theoretically,and the validity of the Hopf bifurcation condition is verified by numerical analysis.The results show that the two time delays can make the stable equilibrium point unstable,thus leading to periodic oscillations induced by Hopf bifurcation.Furthermore,the time delays in FHN and HR neurons have different effects on the firing activity of neural network.Complex firing patterns,such as quiescent state,chaotic spiking,and periodic spiking can be induced by the time delay in FHN neuron,while the neural network only exhibits quiescent state and periodic spiking with the change of the time delay in HR neuron.Especially,phase synchronization between the heterogeneous neurons is explored,and the results show that the time delay in HR neurons has a greater effect on blocking the synchronization than the time delay in FHN neuron.Finally,the theoretical analysis is verified by circuit simulations.

Stability and Hopf bifurcation of a nonlinear electromechanical coupling system with time delay feedback

The stability and the Hopf bifurcation of a nonlinear electromechanical coupling system with time delay feedback are studied. By considering the energy in the air-gap field of the AC motor, the dynamical equation of the electromechanical coupling transmission system is deduced and a time delay feedback is introduced to control the dynamic behaviors of the system. The characteristic roots and the stable regions of time delay are determined by the direct method, and the relationship between the feedback gain and the length summation of stable regions is analyzed. Choosing the time delay as a bifurcation parameter, we find that the Hopf bifurcation occurs when the time delay passes through a critical value.A formula for determining the direction of the Hopf bifurcation and the stability of the bifurcating periodic solutions is given by using the normal form method and the center manifold theorem. Numerical simulations are also performed, which confirm the analytical results.

Hopf Bifurcation and Stability Analysis for a Predator-prey Model with Time-delay

In this paper, a predator-prey model of three species is investigated, the necessary and sucient of the stable equilibrium point for this model is studied. Further, by introduc-ing a delay as a bifurcation parameter, it is found that Hopf bifurcation occurs when τ cross some critical values. And, the stability and direction of hopf bifurcation are determined by applying the normal form theory and center manifold theory. numerical simulation results are given to support the theoretical predictions. At last, the periodic solution of this system is computed.

Hopf Bifurcation of Nonresident Computer Virus Model with Age Structure and Two Delays Effects

This paper constructed and studied a nonresident computer virus model with age structure and two delays effects. The non-negativity and boundedness of the solution of the model have been discussed, and then gave the basic regeneration number, and obtained the conditions for the existence and the stability of the virus-free equilibrium and the computer virus equilibrium. Theoretical analysis shows the conditions under which the model undergoes Hopf bifurcation in three different cases. The numerical examples are provided to demonstrate the theoretical results.

Bifurcation and stability of an improved time-delayed fluid flow model in internet congestion control

Based on the fluid flow time-delayed model proposed by Misra et al in internet congestion control, one modified time-delayed model is presented, where the influence of the communication delay on the router queue length is investigated in detail. The main advantage of the new model is that its stability domain is larger even without an extra controller. By linear stability analysis and numerical simulation, tbe effectiveness and feasibility of the novel model in internet congestion control are verified.

Stability and Hopf bifurcation of a delayed ratio-dependent predator-prey system

Since the ratio-dependent theory reflects the fact that predators must share and compete for food, it is suitable for describing the relationship between predators and their preys and has recently become a very important theory put forward by biologists. In order to investigate the dynamical relationship between predators and their preys, a so-called Michaelis-Menten ratio-dependent predator-prey model is studied in this paper with gestation time delays of predators and preys taken into consideration. The stability of the positive equilibrium is investigated by the Nyquist criteria, and the existence of the local Hopf bifurcation is analyzed by employing the theory of Hopf bifurcation. By means of the center manifold and the normal form theories, explicit formulae are derived to determine the stability, direction and other properties of bifurcating periodic solutions. The above theoretical results are validated by numerical simulations with the help of dynamical software WinPP. The results show that if both the gestation delays are small enough, their sizes will keep stable in the long run, but if the gestation delays of predators are big enough, their sizes will periodically fluc-tuate in the long term. In order to reveal the effects of time delays on the ratio-dependent predator-prey model, a ratiodependent predator-prey model without time delays is considered. By Hurwitz criteria, the local stability of positive equilibrium of this model is investigated. The conditions under which the positive equilibrium is locally asymptotically stable are obtained. By comparing the results with those of the model with time delays, it shows that the dynamical behaviors of ratio-dependent predator-prey model with time delays are more complicated. Under the same conditions, namely, with the same parameters, the stability of positive equilibrium of ratio-dependent predator-prey model would change due to the introduction of gestation time delays for predators and preys. Moreover, with the variation of time delays, the positive equilibrium of the ratio-dependent predator-prey model subjects to Hopf bifurcation.

Stability and Hopf Bifurcation of a Virus Infection Model with a Delayed CTL Immune Response

In this paper, a virus infection model with standard incidence rate and delayed CTL immune response is investigated. By analyzing corresponding characteristic equations,the local stability of each of feasible equilibria and the existence of Hopf bifurcations at the CTL-activated infection equilibrium are established, respectively. By means of comparison arguments, it is verified that the infection-free equilibrium is globally asymptotically stable if the basic reproduction ratio is less than unity. By using suitable Lyapunov functional and LaSalle's invariance principle, it is shown that the CTL-inactivated infection equilibrium of the system is globally asymptotically stable if the immune response reproduction ratio is less than unity and the basic reproduction ratio is greater than unity. Numerical simulations are carried out to illustrate the theoretical result.

Hopf bifurcation analysis of Chen circuit with direct time delay feedback

Direct time delay feedback can make non-chaotic Chen circuit chaotic. The chaotic Chen circuit with direct time delay feedback possesses rich and complex dynamical behaviours. To reach a deep and clear understanding of the dynamics of such circuits described by delay differential equations, Hopf bifurcation in the circuit is analysed using the Hopf bifurcation theory and the central manifold theorem in this paper. Bifurcation points and bifurcation directions are derived in detail, which prove to be consistent with the previous bifurcation diagram. Numerical simulations and experimental results are given to verify the theoretical analysis. Hopf bifurcation analysis can explain and predict the periodical orbit （oscillation） in Chen circuit with direct time delay feedback. Bifurcation boundaries are derived using the Hopf bifurcation analysis, which will be helpful for determining the parameters in the stabilisation of the originally chaotic circuit.

STABILITY AND BIFURCATION OF A HUMAN RESPIRATORY SYSTEM MODEL WITH TIME DELAY

The stability and bifurcation of the trivial solution in the two_dimensional differential equation of a model describing human respiratory system with time delay were investigated. Formulas about the stability of bifurcating periodic solution and the direction of Hopf bifurcation were exhibited by applying the normal form theory and the center manifold theorem.Furthermore,numerical simulation was carried out.

Global Stability and Hopf Bifurcation for a Virus Dynamics Model with General Incidence Rate and Delayed CTL Immune Response

In this work, we investigate an HIV-1 infection model with a general incidence rate and delayed CTL immune response. The model admits three possible equilibria, an infection-free equilibrium E*0, CTL-inactivated infection equilibrium E*1 and CTL-activated infection equilibrium E*2. We prove that in the absence of CTL immune delay, the model has exactly the basic behaviour model, for all positive intracellular delays, the global dynamics are determined by two threshold parameters R0 and R1, if R0 ≤ 1, E*0 is globally asymptotically stable, if R1 ≤ 1 < R0, E*1 is globally asymptotically stable and if R1 >1, E*2 is globally asymptotically stable. But if the CTL immune response delay is different from zero, then the behaviour of the model at E*2 changes completely, although R1 > 1, a Hopf bifurcation at E*2 is established. In the end, we present some numerical simulations.

Bifurcation and Stability Analysis of HIV Infectious Model with Two Time Delays

The HIV problem is studied by version of delay mathematical models which consider the apoptosis of uninfected CD4+ T cells which cultured with infected T cells in big volume. The opportunistic infection and the apoptosis of uninfected CD4+ T cells are caused directly or indirectly by a toxic substance produced from HIV genes. Ubiquitously, the nonlinear incidence rate brings forth the increasing number of infected CD4+ T cells with introduction of small time delay, and in addition, there also exists a natural time delay factor during the process of virus replication. With state feedback control of time delay, the bifurcating periodical oscillating phenomena is induced via Hopf bifurcation. Mathematically, with the geometrical criterion applied in the stability analysis of delay model, the critical threshold of Hopf bifurcation in multiple delay differential equations which satisfy the transversal condition is derived. By applying reduction dimensional method combined with the center manifold theory, the stability of the bifurcating periodical solution is analyzed by the perturbation near Hopf point.

HOPF BIFURCATION OF AN OSCILLATOR WITH QUADRATIC AND CUBIC NONLINEARITIES AND WITH DELAYED VELOCITY FEEDBACK

This paper studies the local dynamics of an SDOF system with quadratic and cubic stiffness terms,and with linear delayed velocity feedback.The analysis indicates that for a sufficiently large velocity feedback gain,the equilibrium of the system may undergo a number of stability switches with an increase of time delay,and then becomes unstable forever.At each critical value of time delay for which the system changes its stability,a generic Hopf bifurcation occurs and a periodic motion emerges in a one-sided neighbourhood of the critical time delay.The method of Fredholm alternative is applied to determine the bifurcating periodic motions and their stability.It stresses on the effect of the system parameters on the stable regions and the amplitudes of the bifurcating periodic solutions.

BIFURCATION ANALYSIS IN A PREDATOR-PREY MODEL WITH AN ALLEE EFFECT AND A DELAYED MECHANISM

Regarding delay-induced predator-prey models, much research has been done on delayed destabilization, but whether delays are stabilizing or destabilizing is a subtle issue. In this study, we investigate predator-prey dynamics affected by both delays and the Allee effect. We analyze the consequences of delays in different feedback mechanisms. The existence of a Hopf bifurcation is studied, and we calculate the value of the delay that leads to the Hopf bifurcation. Furthermore, applying the normal form theory and a center manifold theorem, we consider the direction and stability of the Hopf bifurcation. Finally, we present numerical experiments that validate our theoretical analysis. Interestingly, depending on the chosen delay mechanism, we find that delays are not necessarily destabilizing. The Allee effect generally increases the stability of the equilibrium, and when the Allee effect involves a delay term, the stabilization effect is more pronounced.

A modified averaging scheme with application to the secondary Hopf bifurcation of a delayed van der Pol oscillator

In this paper, a modified averaging scheme is presented for a class of time-delayed vibration systems with slow variables. The new scheme is a combination of the averaging techniques proposed by Hale and by Lehman and Weibel, respectively. The averaged equation obtained from the modified scheme is simple enough but it retains the required information for the local nonlinear dynamics around an equilibrium. As an application of the present method, the delay value for which a secondary Hopf bifurcation occurs is successfully located for a delayed van der Pol oscillator.

Robust exponential stability of uncertain discretetime impulsive switching systems with state delay

A new class of hybrid impulsive and switching models are introduced and their robust exponential stability and control synthesis are addressed. The proposed switched system is composed of stable subsystems and unstable subsystems, which not only involves state delay and norm-bounded time-varying parameter uncertainties, but also contains the impulsive switching effects between the subsystems. Based on the extension of the system dimension and the concept of average dwell time, a kind of practically useful switching rule is presented which guarantees the desired robust exponential stability. A switched state feedback controller is also given.

FLIP AND N-S BIFURCATION BEHAVIOR OF A PREDATOR-PREY MODEL WITH PIECEWISE CONSTANT ARGUMENTS AND TIME DELAY

In this paper, the stability and bifurcation behaviors of a predator-prey model with the piecewise constant arguments and time delay are investigated. Technical approach is fully based on Jury criterion and bifurcation theory. The interesting point is that the model will produce two different branches by limiting branch parameters of different intervals. Besides, image simulation is also given.

一类具有双时滞的捕食模型的稳定性和Hopf分支

建立了一类具有B-D型功能反应和不同避难行为的双时滞捕食模型,并以时滞作为参数,分析了模型在不同时滞情况下正平衡点的稳定性和Hopf分支.研究结果表明,当时滞等于相应的临界值时,模型在正平衡点处发生Hopf分支,且正平衡点的稳定性发生改变.

Tumor-Immune Interaction System with the Effect of Time Delay and Hyperglycemia on the Breast Cancer Cells

Breast cancer in women is a complicated and multifaceted disease. Studies have demonstrated that hyperglycemia is one of the most significant risk factors for breast cancer. Hyperglycemia is when the sugar level in human blood is too high, which means excess glucose. Glucose excess can encourage the growth, invasion, and migration of breast cancer cells at the cellular level. Though, the effects of glucose on the dynamics of breast cancer cells have been examined mathematically by a system of ordinary differential equations. However, the non-instantaneous biological occurrences leading to the secretion of immuno-suppressive cytokines by tumors to evade immune surveillance and the immune cells’ derivation of cytokines to attack the tumor cells are not yet discussed. Therefore, investigating the biological process involved in the dynamics of tumors, immune and normal cells with excessive glucose concentration is inviolable to determining the best procedure for controlling tumors’ uncontrollable growth. Time delay, denoted by τ, is used to describe the time tumor cells take to secrete immunosuppressive cytokines to evade immune surveillance and the time immune cells take to recognize and attack the tumor cells. We have studied the local stability analysis of the biological steady states in both delayed and non-delayed system. The Routh-Hurwitz stability criterion is used to analyze the dynamical equilibrium of the cells’ population. Hopf bifurcation was analyzed by using time delay s as a bifurcation parameter. The analytical results suggest an unstable scenario for a tumor-free equilibrium point as normal cells are bound to grow to their carrying capacity. The result predicts a stable system for coexisting equilibrium when the interaction is instantaneous (τ = 0). However, when τ > 0, the coexisting equilibrium point switches from stable to unstable. The numerical results not only validate all the analytical results but also show the case of possible situations when glucose concentration is varied, indicating that both tumor growth and immune system efficiency are highly affected by the level of glucose in the blood. This concluded that the delay in the secretion of cytokines by immune cells and derivation cytokines by the tumors helps to identify the possible chaotic situation under different glucose concentration and the extent to which such delay can have on restoration of the normal cells when glucose concentration is low.

具有时滞和线性收获项的三维Lotka-Volterra合作系统的Hopf分支

针对三维Lotka-Volterra合作系统的Hopf分支进行了研究。首先,在三维Lotka-Volterra合作系统的基础上,引入时滞项及线性收获项对该系统进行改进;其次,以时滞τ作为分支参数,对改进后系统的局部Hopf分支的存在性进行分析,给出使系统在正平衡点处产生Hopf分支的临界值;然后,利用中心流形定理和规范型理论计算出Hopf分支方向及周期解稳定性的公式;最后,进行数值模拟,根据数值模拟的结果,验证了理论的可行性。结果表明,当时滞τ从0一直增加到超过临界值时,系统的正平衡点由稳定状态变为不稳定状态,且在正平衡点处产生Hopf分支。说明时滞影响了系统的动力学行为,使其动力学性质更加复杂。

一类具有时滞的Leslie-Gower捕食-食饵模型的Hopf分支

利用Hopf分支理论,研究一类具有时滞的Leslie-Gower捕食-食饵模型.首先,以时滞为分支参数,讨论该模型正平衡点的稳定性和Hopf分支的存在性;其次,根据偏泛函微分方程的规范型理论和中心流形定理,确定Hopf分支的分支方向和分支周期解的稳定性;最后,利用MATLAB进行数值模拟.