Variation in photosynthetic photon flux density within a tropical seasonal rain forest of Xishuangbanna, south-western China

The effects of canopy development, solar angle, and weather conditions on temporal variation in photosynthetic photon flux density（PPFD） at three heights within a tropical rain forest canopy in Xishuangbanna, China, were examined. PPFD was measured every second and stored as 10-min averages from 1 December 2002 to 30 November 2003. PPFD variability was examined at three different temporal scales. Specific days in March, September, and December with clear and overcast sky conditions were selected to separate the effects of leaf area index（LAI） and solar angle on diurnal variability. On both clear and overcast days, mean daily average PPFD was significantly different between March and September at all heights, except 10 m on clear days, suggesting that LAI directly influences PPFD. In contrast, the differences in daily average PPFD among three heights between September and December were likely due to variation in solar angle. In addition, daily average PPFD at all locations were significantly lower under overcast than clear sky conditions in March, September and December. Over the year-long study, the mean daily total PPFD at 2! m, 10 m and 4 m was 2.8, 2.7 and 0.7 mol/（m^2·d）, which accounted for 9.7%, 9.4% and 2.4% of the daily PPFD above the canopy, respectively. Significant differences in mean daily total PPFD occurred at the same heights among different seasons, and diurnal, day-to-day and seasonal PPFD varied at different heights within the canopy. The possible effects of light variability on physiological and morphological responses of plants are discussed.

Risk Assessment Mapping of Landscape Development Based on Ecological Service and Goods in Malaysia Lowland Tropical Rainforest

Amongst the impacts of converting forest to agricultural activities are soil erosion and degradation of ecology service values and goods （ESVG）. The soil erosion can be seen as on-site impacts, such as the problems of decreasing soil fertility and also its off-site impact such as the problems of sedimentation of the nearby rivers, whilst the degradation of ESVG are more holistie in nature, These impacts can be devastating in environmental, biological, and socio-economic manners. This paper reports the study undertaken on the impacts of agricultural development in 0.8 million ha of forest dominated landscape in Pasoh Forest Region （PFR）, Malaysia, within period of 8 years from 1995 to 2003. Three folds of impacts on agricultural development examined and analysed, are： （i） relationship of total soil loss and changes in land use pattern, （ii） mapping trends of ESVG for PFR in 1995 and 2003, and （iii） risk assessment of ESVG based on simulation of converting 339,630 ha of primary forest into mass-scale oil palm plantation. Results of this study indicated that although only minor changes of about 1464 ha （about 0.2% of PFR） of primary forest was converted to agricultural activities, it have significantly increased the total soil loss from 59 to 69 million ton/ha/yr. The mean rate of soil is loss for PFR is 0.8 mil ton/ha/yr and if translated into ESVG term, the soil loss costs about US$ 4.8mil/yr. However, majority of the soil loss within all land use classes are within range of very low-low risk categories （〈10 ton/ha/yr）. ESVG for PFR were costing US$ 179 millions in 1995, declined to US$114 millions in 2003 due to 0.2% reduction of forested land. The ESVG of converting 339,630 ha primary forest into mass plantation cost less than original forest within period of 20 years examined; the 20th year of conversion, the ESVG of plantation and to-remain as forest cost US$ 963 and US$ 575 millions, respectively. However, this difference is only marginal when full attributes of ESVG are considered.

Growth Traits and the Trade-Offs for Tree Species with Arbuscular Mycorrhizal Fungi in a Tropical Rain Forest Edge at Los Tuxtlas,Mexico

The effect of arbuscular mycorrhizal fungi on seedling growth across the rain forest-pasture edge has not received much attention. In a tropical rain forest in eastern Mexico, the seedlings of light demanding (Ficus insipida), nonsecondary light demanding (Lonchocarpus cruentus) and shade tolerant species (Nectandra ambigens, Coccoloba hondurensis) were grown and transplanted to a forest edge with three inoculation treatments (AM fungus spores and colonized roots, spores, and no inoculum). For all species, stem height, stem diameter, total dry weight, leaf area and net assimilation rate were higher in the pasture. Stem height, stem diameter and root/shoot were higher for L. cruentus, and leaf area ratio, specific leaf area and net assimilation rate were higher for F. insipida;the lowest values of almost all variables were recorded for N. ambigens. L. cruentus and C. hondurensis with mycorrhizae had the highest values for root/shoot and net assimilation rate, respectively. The lowest values of root/shoot and net assimilation rate were observed for nonlight-demanding species in the forest. There were clear trade-offs for the pioneer species between survival and growth, and in underground biomass allocation and assimilation for nonsecondary light demanding, but there was not for the shade-tolerant species.

PROTECTING THE RESOURCES OF THE TROPICAL RAIN FORESTS

In order to finish the cataloging work of buttercup species in the family of Renunculaceae for the compilation of the Flora Yunnanensis, I went to the CAS Institute of Botany in Kunming City,capital of Yunnan Province in May 1995 to look up phyto-taxonomic specimens. At the invitation of the Institute’s director Prof. Xu Zhaifu, when the work was coming to an end, I toured the well-known botanic garden of the Institute in the township of Menglu, which nests in the picturesque district of Xishuang Banna at the southernmost tip of the subtropical territory of Yunnan. In the winter of 1958,I had visited the ever-verdant vicinity to collect floral materials. After almost 37 years, I revisited the botanic garden founded by celebrated

Fragmented forest in tropical landscape——the case of the State of Selangor, Malaysia

This paper presents the pattern and changes of fragmented forest in relation with changes of total forest cover in the state of Selangor in three decades. In this study, inventoried forest cover maps of Selangor in 1971/1972, 1981/1982 and 1991/1992 produced by the Forestry Department of Peninsular Malaysia were digitized to examine the changes in area and number of fragmented forest. Results showed that in 1971/1972, 16 fragmented forests were identified in Selangor. All fragmented forests were identified as dipterocarp forest. A decade later the number of fragmented forests increased by approximately 44% (23). Of the 23 fragmented forests, two were peat swamp forests whereas the remaining were dipterocarp forests. In 1991/1992 the number of fragmented forests (12) was reduced by 47.8%. Two of the fragmented forests were identified as peat swamp forest, seven dipterocarp forest and the other three was mixed of dipterocarp forests and plantation forests. Fragmentation of both dipterocarp and peat swamp forests occurred profoundly during the period between 1971/1972 and 1981/1982, which consequently increased the number of fragmented forests compared with before the period of 1971/1972 where fragmentation happened only at dipterocarp forests. However, many fragmented forests vanished between the 1981/1982 and 1991/1992 periods.

Old-growth mixed dipterocarp forests show variable losses and gains in aboveground biomass and standing carbon over forty years

Background:No studies have documented long-term trends in aboveground biomass(AGB)for mixed-dipterocarp forests(MDF),the dominant rain forest type in tropical wet equatorial Asia.In our study,we sought to document such trends over forty years across three sites representing lowland to lower montane elevations.Methods:To do this,we established fifty 100 m×25 m plots in 1978 across three sites sampled along an elevation gradient,identified as mature old-growth forest.We measured trees for diameter at breast height that we identified to species and tagged.We took wood samples to calculate species wood-specific gravity.We re-measured plots in 1998 and again in 2018.Results:We show standing AGB for all sites combined to be 517.52 Mg·ha^(-1)in 1978,but this declined by 17%over 40 years to 430.11 Mg·ha^(-1).No differences exist among sites in AGB primarily because of considerable within site variation;but interactions of time with site show declines across sites were not uniform,one remained about the same.Relatively few species represented a high proportion of the AGB with the top five species comprising between 34%and 65%,depending upon site and year sampled.One species,Mesua nagassarium,represented a disproportionately large amount of AGB and decline over time,particularly at the low elevation site.Conclusions:Our results are directly relevant to estimating AGB and standing carbon sequestered in MDF.Our study is the first to demonstrate varying but overall,declining trends in amounts of AGB among forests making predictions of biomass and standing carbon in MDF difficult over wide regions.

Forest vegetation of Xishuangbanna,south China

Xishuangbanna of southern Yunnan is biogeographically located at a transitional zone from tropical southeast （SE） Asia to subtropical east Asia and is at the junction of the Indian and Burmese plates of Gondwana and the Eurasian plate of Laurasia. The region, though surprisingly far from the equator and at a relatively high altitude, has a rich tropical flora and a typical tropical rain forest in the lowland areas. Based on physiognomic and ecological characteristics, floristic composition and habitats combined, the primary vegetation in Xishuangbanna can be organized into four main vegetation types： tropical rain forest, tropical seasonal moist forest, tropical montane evergreen broad-leaved forest and tropical monsoon forest. The tropical rain forest can be classified into two subtypes, i.e. a tropical seasonal rain forest in the lowlands and a tropical montane rain forest at higher elevations. The tropical seasonal rain forest has almost the same forest profile and physiognomic characteristics as equatorial lowland rain forests and is a type of truly tropical rain forest. Because of conspicuous similarity on ecological and floristic characteristics, the tropical rain forest in Xishuangbanna is a type of tropical Asian rain forest. However, since the tropical rain forest of Xishuangbanna occurs at the northern edge of tropical SE Asia, it differs from typical lowland rain forests in equatorial areas in having some deciduous trees in the canopy layer, fewer megaphanerophytes and epiphytes but more abundant lianas and more plants with microphyll. It is a type of semi-evergreen rain forest at the northern edge of the tropical zone. The tropical montane rain forest occurs at wet montane habitats and is similar to the lower montane rain forest in equatorial Asia in floristic composition and physiognomy. It is a type of lower montane rain forests within the broader category of tropical rain forests. The tropical seasonal moist forest occurs on middle and upper limestone slopes. It is similar to the tropical montane evergreen broad-leaved forest in the region in physiognomy, but differs from the latter in floristic composition. It is a vegetation type on limestone at high elevations. The monsoon forest in Xishuangbanna is a tropical deciduous forest under the influence of a strong monsoon climate and is considered to be a transitional vegetation type between a tropical rain forest and savanna in physiognomy and distribution. The tropical montane evergreen broad-leaved forest is the main montane vegetation type in the region. It is dominated largely by the families Fagaceae, Euphorbiaceae, Theaceae and Lauraceae. It differs from tropical lower montane rain forests in its lack of epiphytes and in having more abundant lianas and plants with compound leaves. It is considered to be a distinct vegetation type from the northern margin of mainland southeastern Asia, controlled by a strong seasonal climate, based on its floristic and physiognomic characteristics.

Spatial and seasonal variation in soil respiration along a slope in a rubber plantation and a natural forest in Xishuangbanna, Southwest China

Soil respiration is a key component of the global carbon cycle, and even small changes in soil respiration rates could result in significant changes in atmospheric CO_2 levels. The conversion of tropical forests to rubber plantations in SE Asia is increasingly common, and there is a need to understand the impacts of this land-use change on soil respiration in order to revise CO_2 budget calculations. This study focused on the spatial variability of soil respiration along a slope in a natural tropical rainforest and a terraced rubber plantation in Xishuangbanna, Southwest(SW) China. In each land-use type, we inserted 105 collars for soil respiration measurements.Research was conducted over one year in Xishuangbanna during May, June, July and October 2015(wet season) and January and March 2016(dry season). The mean annual soil respiration rate was 30% higher in natural forest than in rubber plantation and mean fluxes in the wet and dry season were 15.1 and 9.5 Mg C ha^(-1) yr^(-1) in natural forest and 11.7 and 5.7 Mg C ha^(-1) yr^(-1) in rubber plantation. Using a linear mixedeffects model to assess the effect of changes in soil temperature and moisture on soil respiration, we found that soil temperature was the main driver of variation in soil respiration, explaining 48% of its seasonal variation in rubber plantation and 30% in natural forest. After including soil moisture, the model explained 70% of the variation in soil respiration in natural forest and 76% in rubber plantation. In the natural forest slope position had a significant effect on soil respiration, and soil temperature and soil moisture gradients only partly explained this correlation. In contrast, soil respiration in rubber plantation was not affected by slope position, which may be due to the terrace structure that resulted in more homogeneous environmental conditions along the slope. Further research is needed to determine whether or not these findings hold true at a landscape level.

Hydrological Processes in a Small Research Watershed under Forest Coverage in the Coast of Chiapas, Mexico

In the hydrological watershed, some natural processes take place in which the interaction of water, soil, climate and vegetation favors the capture of water. The present study aimed to evaluate preliminary information regarding the hydrological response and the water balance in a small research watershed with tropical forest cover (15°01'44''N and 92°13'55''W, 471 m, 2.3 has). Events of precipitation, direct runoff, infiltration rate and baseflow were performed. The amount, duration and intensity of rainfall events were recorded with the use of a pluviograph. Surface runoff was quantified with an established gauging station, an H-type gauging device and a horizontal mechanical gauging limnograph. Runoff base flow was measured at the gauging station using the volume-time method. Infiltration was measured using a triple ring infiltrometer, taking two measurements in the upper part and two in the lower part of the microbasin. Evapotranspiration was measured with the amount of rainfall entering and runoff leaving the watershed. In the study period, annual rainfall of 4417.6 mm distributed over 181 events were recorded;about 70% of the storms showed lower intensities at 20 mm·h-1. The total runoff was 345.8 mm caused by half of the rainfall events, which represents 7.8% of the total rain;77% of runoff events showed lower sheets of 5 mm and an average specific rate of 20.7 L·s-1·ha-1 with a maximum of 113.6 L·s-1·ha-1. Three runoff events were greater than 20.1 mm and caused the 22.5% of the total runoff depth in the study period showing the equilibrium conditions in the hydrological response of the forest. Water outputs like baseflow was 669.5 mm. In this way, 90% of the rainfall is infiltrated every year in the micro-watershed, which shows the importance of the plant cover in the hydrological regulation and the groundwater recharge.

The effect of water-level fluctuations on swamp forest colonization by seedlings of <i>Tabebuia cassinoides</i>DC. (Bignoniaceae)

Establishment of Tabebuia cassinoides seedlings is related to water-level fluctuations in southeastern Brazil swamp forests. Nine years of annual monitoring of 48 individuals established during a drought in November 1997, when the swamp was unflooded, suggested that their establishment depends on this unpredictable event. This conclusion is further sustained by the wide variability of the seedling cohort size structure, and the fascicular root conformation that holds the shoots erect.

Biomass and carbon dynamics of a tropical mountain rain forest in China

Biometric inventories for 25 years,from 1983 to 2005,indicated that the Jianfengling tropical mountain rain forest in Hainan,China,was either a source or a modest sink of carbon.Overall,this forest was a small carbon sink with an accumulation rate of(0.56±0.22) Mg C ha-1yr-1,integrated from the long-term measurement data of two plots(P9201 and P8302).These findings were similar to those for African and American rain forests((0.62±0.23) Mg C ha-1yr-1).The carbon density varied between(201.43±29.38) Mg C ha-1 and(229.16±39.2) Mg C ha-1,and averaged(214.17±32.42) Mg C ha-1 for plot P9201.Plot P8302,however,varied between(223.95±45.92) Mg C ha-1 and(254.85±48.86) Mg C ha-1,and averaged(243.35±47.64) Mg C ha-1.Quadratic relationships were found between the strength of carbon sequestration and heavy rainstorms and dry months.Precipitation and evapotranspiration are two major factors controlling carbon sequestration in the tropical mountain rain forest.

Annual variation of carbon flux and impact factors in the tropical seasonal rain forest of Xishuangbanna, SW China

Two years of eddy covariance measurements of above- and below-canopy carbon fluxes and static opaque chamber and gas chromatography technique measurements of soil respiration for three treatments (bare soil, soil+litterfall, soil+litterfall+seedling) were carried out in a tropical seasonal rain forest. In addition, data of photosynthesis of dominant tree species and seedlings, leaf area index, litter production and decomposing speed, soil moisture, soil temperature and photosynthetic photon flux density within the forest were all measured concurrently. Data from January 2003 to December 2004 are used to present annual variability of carbon flux and relationships between carbon flux and impact factors. The results show that carbon flux of this forest presented unusual tendency of annual variation; above-canopy carbon fluxes were negative in the dry season (November-April) and mainly positive in the rainy season, but overall the forest is a carbon sink. Carbon flux has obviously diurnal variation in this tropical seasonal rain forest. Above-canopy carbon fluxes were negative in the daytime and absolute values were larger in the dry season than that in the rainy season, causing the forest to act as a carbon sink; at night, carbon fluxes were mainly positive, causing the forest to act as a carbon source. Dominant tree species have greater photosynthesis capability than that of seedlings, which have a great effect on above-canopy carbon flux. There was a significant correlation between above-canopy carbon flux and rate of photosynthesis of tree species. There was also a significant correlation between above-canopy carbon flux and rate of photosynthesis of seedlings; however, the below-canopy carbon flux was only significantly correlated with rate of photosynthesis of seedlings during the hot-dry season. Soil respiration of the three treatments displayed a markedly seasonal dynamic; in addition, above-canopy carbon fluxes correlated well with soil respiration, litterfall production, litterfall decomposition rate, precipitation, and soil moisture and temperature. A primary statistical result of this study showed that above-canopy carbon flux in this forest presented carbon source or sink effects in different seasons, and it is a carbon sink at the scale of a year.

海南热带雨林国家公园森林分类经营实践研究

以海南热带雨林国家公园森林资源分类管理实例为基础,经过实地考察调研,确定了生物及林木资源现状。研究结果表明:海南热带雨林国家公园内的森林类型可划分为严格保护经营类型组、保育修复经营类型组、人工促进修复经营类型组、限制性经营类型组和科普游憩经营类型组5个类型组;依照森林资源实际及森林经营类型,制定了5种森林经营作业法,并采取“封山育林为主,生态修复为辅,注重生物多样性保护”等保护措施对海南热带雨林国家公园中的森林进行保护。研究结果可为热带雨林中的森林分类经营提供实践经验和技术支持。

A preliminary study on the heat storage fluxes of a tropical seasonal rain forest in Xishuangbanna

In order to discuss the values and daily variation characteristics of heat storage fluxes in a tropical seasonal rain forest in Xishuangbanna, the sensible and latent heat storage flux within air column, canopy heat storage flux, energy storage by photosynthesis and ground heat storage above the soil heat flux plate, as well as the ratios of these heat storage fluxes to the net radiation in the cool-dry, hot-dry and rainy season were compared and analyzed based on the observation data of carbon fluxes, meteorological factors and biomass within this tropical seasonal rain forest from January 2003 to December 2004. The findings showed that heat storage terms ranged significantly in the daytime and weakly in the nighttime, and the absolute values of sensible and latent heat storage fluxes were obviously greater than other heat storage terms in all seasons. In addition, the absolute values of total heat storage fluxes reached the peak in the hot-dry season, then were higher in the rainy season, and reached the minimum in the cool-dry season. The ratios of heat storage fluxes to net radiation generally decreased with time in the daytime, moreover, the sensible and latent heat storage dominated a considerable fraction of net radiation, while other heat storage contents occupied a smaller fraction of the net radiation and the peak value was not above 3.5%. In the daytime, the ratios of the total heat storage to net radiation were greater and differences in these ratios were distinct among seasons before 12:00, and then they became lower and differences were small among seasons after 12:00. The energy closure was improved when the storage terms were considered in the energy balance, which indicated that heat storage terms should not been neglected. The energy closure of tropical seasonal rain forest was not very well due to effects of many factors. The results would help us to further understand energy transfer and mass exchange between tropical forest and atmosphere. Moreover, they would supply a research basis for studying energy closure at other places.

Seed Dynamics in Relation to Gaps in a Tropical Montane Rainforest of Hainan Island,South China:(Ⅰ)Seed Rain

Seed dynamics is an important part of stand dynamics in forest ecosystems. In this paper, 26 gaps were randomly selected to study the influence of gaps on the spatial and temporal patterns of seed rains in a tropical montane rainforest of Hainan Island, South China. Three zones for each gap, including outside gap zone （Non-gap）, transitional gap zone （EG-CG）, and central gap zone （CG）, were designed, and fourseed traps （each lm x lm in size） were placed in each zone. Seed rains were collected by these traps every 10 days from June 2001 to May 2002. Seed rain varied greatly with season and generally exhibited a pattern of unimodal change during the study period： seed abundance and species richness were both greater in the wet season than in the dry season. Gaps significantly influenced the temporal patterns of both species richness and density of seed rains. Gaps had no significant influences on the spatial distribution patterns of seed rain species richness, but significantly affected the spatial distribution pattern of seed rain densities. Among the three different zones of gaps, the outside gap zone generally received more seeds inputs than the two other gap zones.

Soil respiration and its partitioning in different components in tropical primary and secondary mountain rain forests in Hainan Island,china

Aims Soil respiration is one of the most important components in the car-bon(c)cycle in terrestrial ecosystems.to investigate the contribution of each component of c cycle to the total soil c efflux,we quantified the rates of litter,root,and other mineral soil respiration from 2012 to 2014 in the primary and secondary tropical mountain rain forests in Hainan Island,china.Methods the seasonal dynamics of soil(Rs),non-litter(RNL)and non-root(RNR)respiration rates were measured using an automatic chamber system(Li-8100).Litter removal and root removal treatments were used to assess the contribution of litter and roots to belowground c production.We estimated the annual c efflux of each compo-nent of soil respiration in primary and secondary forests using a temperature-based exponential model and analyzed the impact of each component in each forest type.Important Findingsthe annual total soil c efflux was significantly higher in the primary rain forest(1567±205 g c m^(−2)yr^(−1))than that in the secondary forest(1300±70 g c m^(−2)yr^(−1),P<0.05).the litter,root,and mineral soils contributed 22%(349±185 g c m^(−2)yr^(−1)),38%(589±100 g c m^(−2)yr^(−1)),and 40%(628±128 g c m^(−2)yr^(−1))to the total soil c efflux in primary rain forest,respectively.In secondary forest,these three components contributed 11%(148±35 g c m^(−2)yr^(−1)),45%(572±259 g c m^(−2)yr^(−1)),and 44%(580±226 g c m^(−2)yr^(−1)),respectively.the temperature sensitivity(Q10)of Rs(2.70±0.14)in the primary forest was significantly higher than that in the secondary forest(2.34±0.12),with the Q10 values for respiration decreasing in the order of RNR>Rs>RNL.these results show that the difference in litter respiration between primary and secondary forest caused the major difference in annual soil respiration efflux between these two forest types.In addition,the litter respiration is more sensitive to the soil temperature than the other soil respiration components.

Gap Dynamics and Tree Species Diversity in a Tropical Montane Rain Forest of Hainan Island,China

Based on investigation of 53 gaps and 25 quadrats (15m×15m each) of non-gap closed stand in an old-growth tropical montane rain forest of Hainan Island, China, canopy disturbance regimes and gap regeneration were studied. Gaps were elliptical in horizontal form, the ratio of long axis /short axis was about 1.4. Percentage of expanded gaps (EG) and canopy gaps (CG) area in the landscape were 53.5% and 25.2% respectively. EG ranged from 31.4 m2 to 488.2m2 and CG/rom 14.9m2 to 354.2m2, their average sizes ...

建设森林防火设施对国家公园生物多样性的影响

通过现场调查与查阅文献相结合的方式,依据相关评价技术规范,评价建设森林防火设施对海南热带雨林国家公园生物多样性的影响。结果显示,建设森林防火设施对国家公园生物多样性影响指数为50.2,属于中低度影响。森林火灾高风险区综合治理建设对海南热带雨林国家公园生物多样性影响程度低。

西双版纳热带季节雨林生态系统氮的生物地球化学循环研究

用小流域集水区和物质平衡方法 ,于 1999年对西双版纳热带季节雨林生态系统的氮素循环进行了初步研究。西双版纳季节雨林生态系统的氮库总储量为 6 4 81.2kg·hm-2 ,其中活体生物量、凋落物层和土壤 (0～ 30cm)中的氮储量分别为 970 .9、37.7、5 4 81.2kg·hm-2 。土壤中的氮占生态系统氮总储量的 84 .4 % ,活体生物量占 15 .0 % ,凋落物层仅占 0 .6 %。结果表明季节雨林的氮主要分布在土壤中 ,而在生物量中只占很少部分。大气降水、林内穿透水、树干流及地表径流的氮含量分别为 0 .5 6 5、0 .82 8、0 .983和 1.0 4 2mg·dm-3 ,氮通量则分别为 8.89、10 .97、3.5 7、5 .95kg·hm-2 ·a-1。大气降水输入氮 8.89kg·hm-2 ·a-1,径流输出氮 5 .95kg·hm-2 ·a-1,收支平衡 (输入—输出 )为2 .94kg·hm-2 ·a-1。氮的生物循环 :吸收为 14 9.86kg·hm-2 ·a-1,存留为 6 9.30kg·hm-2 ·a-1,归还为 80 .5 6kg·hm-2 ·a-1,循环系数为 0 .5 4。结果表明未受干扰的季节雨林生态系统处于氮积累的状态 ,有利于该生态系统的稳定与持续发展。

西双版纳不同海拔热带雨林凋落量变化研究

在3个海拔梯度(600、1 100和1 600 m)选取8块热带雨林样地,研究了印度季风环境下西双版纳热带季节雨林和山地雨林的凋落动态随海拔的变化及其与气候的关系.在3个梯度上,年平均温度分别为22.1、20.1和16.6℃,年平均温度递减率为-0.005 3℃·m-1.随海拔增加,年平均降雨量增加(分别为1 532、1 659和2 011 mm),但旱季的降雨量基本相同(282～295 mm);年蒸发量变化较小(分别为1 369、1 374和1 330mm);年平均空气相对湿度降低(分别为86%、81%和84%),旱季后期湿度降低更明显;样地土壤含水显著增加.热带季节雨林凋落量(1 072～1 285 g·m-2·a-1)显著高于热带山地雨林凋落量(718～1 014 g·m-2·a-1).凋落量和凋落进程变异系数与海拔之间存在线性显著负相关,凋落量与温度线性显著正相关而与降雨量显著负相关.旱季凋落高峰受到空气相对湿度和土壤含水量影响,随海拔增加空气相对湿度降低使得海拔1 105～1 720 m的凋落高峰提前,但土壤含水量继续增加又会使凋落高峰推后.研究结果得出:1)热带季节雨林凋落量与东南亚热带潮湿雨林相近;2)旱季水分限制随海拔增加而变化,影响凋落高峰出现时间;3)随海拔增加,热带山地雨林凋落年进程由季节性向平稳过渡.