基金Supported by the Beijing Nova Program(2008B37)the Major Research Program of Beijing Municipal Commission of Science and Technology(D131100000413001)+2 种基金National Key Technologies R&D Program(2012BAK26B03)the Special Fund for Construction of Scientific and Technological Innovation Ability of Beijing Academy of Agriculture and Forestry Sciencesthe Special Research Fund for the Youth of Beijing Academy of Agriculture and Forestry Sciences(QNJJ201211)~~