基金Acknowledgements This work was supported by the National Natural Sciences Foundation of China (NSFC 31272268/ 31172113), the Special Foundation of Ministry of Science and Technology of the People' s Republic of China (MOST grant no. 2014FY110100) and the Natural Science Foundation of Guizhou Province (J [2014] 2146). We are grateful to, Jianyong Li, Ziqi Li, Yaqian Fu and, Mu Yan, for collecting specimens. This work was supported by the National Natural Sciences Foundation of China (NSFC-31272268/31172113), the Special Foundation of Ministry of Science and Technology of the People's Republic of China (MOST grant no. 2014FY 110100) and the Natural Science Foundation of Guizhou Province (J [2014] 2146).