基金the National Natural Science Foundation of China(Grant No.31000947)the Open Fund of Key Laboratory of Freshwater Ecology and Healthy Aquaculture,Chinese Academy of Fishery Science(Grant No.2010FEA03016)+1 种基金the Science-Technology Foundation for Middle-aged and Young Scholars of Education Department,Hubei Province(Grant No.Q20101702)Science&Technology Pillar Program of Huaian(Grant No.SN 1087)