基金This study was supported by National Key Research and Development Program of China (2016YFA0601801), the State Key Program of National Natural Science Foundation of China (41530424), National Program on Global Change and Air-Sea Interactions, State Oceanic Administration (SOA) (GASI-IPOVAI-03), and the National Natural Science Foundation of China (41305121). We sincerely thank two anonymous reviewers whose comments improved the paper.