A Review of the Tetrapod Track Record in China,with Special Reference to Type Ichnospecies:Implications for Ichnotaxonomy and Paleobiology

＂Splitting＂ and ＂lumping＂ are perpetual problems in vertebrate, especially dinosaur, ichnotaxonomy. Chinese dinosaur ichnotaxonomy, which began in 1940, provides a series of interesting case studies, highlighting the dual problems of historical and dubious ichnotaxonomy. Chinese Mesozoic tetrapod track types have been placed into 63 ichnospecies （one Triassic, 28 Jurassic, and 34 Cretaceous）, exclusive of other, non-type ichnospecies or ichnotaxa identified from China. Fifty-two （~83%） of these 63 tetrapod ichnospecies were placed in monospecific ichnogenera. At the ichnogenus level, we prune---either by recognizing nomina dubia or by synonymy--17 from the list of 53 dinosaurian ichnogenera （a 32% reduction）, leaving 36 ichnotaxa that we consider valid. Most of the cuts affect Jurassic theropod ichnotaxa, which are reduced from 23 to only nine because most ichnogenera are subjective junior synonyms of Grallator and Eubrontes. Fewer Chinese Cretaceous ichnotaxa （only six of 21 ichnogenera） are obvious nomina dubia or subjective synonyms, suggesting greater east Asian endemism during this time. Because ichnospecies differences are subtle, we provisionally retain ichnospecies as valid pending detailed comparative analyses of congeneric ichnospecies. This synthesis is long overdue and is necessary to address problems of historical and provincial ichnotaxonomy, which severely hamper comparisons of tetrapod ichnofaunas in space and time.

An Unusual, Three-Dimensionally Preserved, Large Hadrosauriform Pes Track from "Mid"-Cretaceous Jiaguan Formation of Chongqing, China

Three-dimensional tracks provide unique insights into the locomotor mechanics of their track makers. An isolated, large hadrosauriform print attributable to Caririchnium lotus from the ＂mid＂-Cretaceous Lotus track site （Jiaguan Formation） in China permits reconstruction of the footfall, weight-bearing, and kick-off phases of the step cycle. Large-scale modifications of the pes during the step cycle indicate C. lotus trackmakers were capable of locomotory modifications in response to substrate consistency beyond the ＂expected＂ shift between bipedal and quadrupedal postures. An unusual curvature to the trace of one of the outer digits indicates substantial transverse mobility. The remaining digits demonstrate lesser degrees of transverse movement accompanied by extension of the digits during footfall. The absence of overprinted scale-scratch marks and toe drags are consistent with a vertical kick-off of the pes and concomitant flexion of the digits. This track suggests that pedal mobility in C. lotus track makers was greater than previously suspected and has implications for reconstructions of hadrosauriform locomotion.

A Large Therizinosauroid(Dinosauria:Theropoda) from the Early Cretaceous of Northwestern China

We herein describe an associated partial postcranial skeleton of a theropod dinosaur discovered in the Lower Cretaceous Xinminpu Group of the Yujingzi Basin, in the Jiuquan area of Gausu Province, northwestern China. Features of its humerus, such as strongly expanded proximal and distal ends, a well developed medial tuberosity, distal condyles expressed on the humeral cranial surface, and a hypertrophied entepicondyle, definitively establish the therizinosauroid affinities of the specimen. It differs from other therizinosauroids in having a shallow, poorly demarcated glenoid fossa with a prominent rounded and striated tumescence on the dorsomedial surface of its scapular portion, and a pubis with a strongly concave cranial margin. It represents a new taxon, Suzhousaurus megatherioides gen. et sp. nov. Cladistic analysis recovers Suzhousaurus as the sister taxon of Nothronychus mckinleyi from the mid-Cretaceous of western North America; together, they are basal members of the Therizinosauroidea, more derived than the Early Cretaceous Falcarius and Beipiaosaurus but less derived than Alxasaurus and the Therizinosauridae. Along with ＂Nanshiungosaurus＂ bohlini from possibly coeval beds in the Mazongshan area of northern-most Gansu, Suzhousaurus represents one of the largest-known Early Cretaceous therizinosauroids, demonstrating that this clade attained considerable body size early in its evolutionary history.