基金supported by National Basic Research Program of China (Grant No.2009CB219508)Chinese Academy of Sciences (Grant No.KZCX2-YW-GJ03)+2 种基金National Natural Science Foundation of China (GrantNos. 91228206 & 40725011)GIGCAS 135 Program (Grant No.Y234021001)Scientific and Technological Program of Guangdong Province (Grant No. 2011A080403021)