基金the State Key Laboratory of Palaeobiology and Stratigraphy(Nanjing institute of Geology and Palaeontology,CAS)(Grant No.053109)the National Natural Sciences Foundation of China(Grant Nos.40162002.40372023.40232020)+1 种基金the Foundation of the Key and Basic Project of Science and Technology of Guizhou(Grant No.2002-309) the Earlyand Special Project of Key Basic Project of the Ministry of Science andTechnology.China(Grant No.2002,CCC02600).